{"title":"Orthoceratoid and coleoid cephalopods from the Middle Triassic of Switzerland with an updated taxonomic framework for Triassic Orthoceratoidea","authors":"Alexander Pohle, Christian Klug","doi":"10.1186/s13358-024-00307-8","DOIUrl":"https://doi.org/10.1186/s13358-024-00307-8","url":null,"abstract":"Orthoconic cephalopods are subordinate, but persistent, widespread and regionally abundant components of Triassic marine ecosystems. Here, we describe unpublished specimens from the Anisian (Middle Triassic) Besano Formation at Monte San Giorgio, Switzerland. They can be assigned to two major but unrelated lineages, the Coleoidea and the Orthoceratoidea. The orthoceratoids belong to Trematoceras elegans (Münster, 1841) and occur regularly within the Besano Formation, are uniform in size, and have few available morphological characters. In contrast, coleoids are more diverse and appear to be restricted to shorter intervals. A new coleoid is described as Ticinoteuthis chuchichaeschtli gen. et sp. nov. To better put the orthoceratoids of the Besano Formation into perspective, we also synthesise the current taxonomy of Triassic orthoceratoids on a global scale. The currently used scheme is largely outdated, with very little taxonomic progress in the past 100 years. Despite previous research showing the distinctness of Triassic orthoceratoids from Palaeozoic taxa, they are still commonly labelled as “Orthoceras” or “Michelinoceras”, which are confined to the Palaeozoic. We show that Triassic orthoceratoids probably belong to a single lineage, the Trematoceratidae, which can be assigned to the Pseudorthocerida based on the embryonic shell and endosiphuncular deposits. Many Triassic species can probably be assigned to Trematoceras, but there are at least two additional Triassic orthoceratoid genera, Paratrematoceras and Pseudotemperoceras. Finally, we review the palaeobiogeographic and stratigraphic distribution of the group and outline possible future research directions.","PeriodicalId":56059,"journal":{"name":"Swiss Journal of Palaeontology","volume":null,"pages":null},"PeriodicalIF":3.0,"publicationDate":"2024-04-03","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"140588395","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Stephan N. F. Spiekman, Martín D. Ezcurra, Adam Rytel, Wei Wang, Eudald Mujal, Michael Buchwitz, Rainer R. Schoch
{"title":"A redescription of Trachelosaurus fischeri from the Buntsandstein (Middle Triassic) of Bernburg, Germany: the first European Dinocephalosaurus-like marine reptile and its systematic implications for long-necked early archosauromorphs","authors":"Stephan N. F. Spiekman, Martín D. Ezcurra, Adam Rytel, Wei Wang, Eudald Mujal, Michael Buchwitz, Rainer R. Schoch","doi":"10.1186/s13358-024-00309-6","DOIUrl":"https://doi.org/10.1186/s13358-024-00309-6","url":null,"abstract":"Some of the earliest members of the archosaur-lineage (i.e., non-archosauriform archosauromorphs) are characterised by an extremely elongated neck. Recent fossil discoveries from the Guanling Formation (Middle Triassic) of southern China have revealed a dramatic increase in the known ecomorphological diversity of these extremely long-necked archosauromorphs, including the fully marine and viviparous Dinocephalosaurus orientalis. These recent discoveries merit a reinvestigation of enigmatic Triassic diapsid fossils from contemporaneous European deposits housed in historical collections. Here, we provide a redescription of Trachelosaurus fischeri, represented by a single, disarticulated specimen first described in 1918. Due to its unique morphology, which includes short, bifurcating cervical ribs, and a high presacral vertebral count, this taxon has been referred to either as a “protorosaurian” archosauromorph or a sauropterygian. Our revision clearly shows that Trachelosaurus represents the first unambiguous Dinocephalosaurus-like archosauromorph known from outside the Guanling Formation. Our finding has important systematic implications. Trachelosauridae Abel, 1919 represents the senior synonym for the recently identified Dinocephalosauridae Spiekman, Fraser and Scheyer, 2021. Based on our phylogenetic analyses, which employ two extensive datasets, we also corroborate previous findings that tanystropheids and trachelosaurids represent two families within a larger monophyletic group among non-crocopodan archosauromorphs, which is here named Tanysauria (clade nov.). Trachelosauridae is minimally composed of Trachelosaurus fischeri, Dinocephalosaurus orientalis, Pectodens zhenyuensis, and Austronaga minuta, but one of our analyses also found a probably taxonomically broader clade that may also include Gracilicollum latens and Fuyuansaurus acutirostris. Trachelosaurus fischeri considerably expands the known spatial and temporal range of Trachelosauridae to the earliest Anisian and the Central European Basin. Our findings add to the growing evidence for the presence of a diverse group of fully marine reptiles during the Middle Triassic among Tanysauria. These trachelosaurids possess flipper-like limbs, high vertebral counts, and elongate necks, thus superficially resembling long-necked Jurassic and Cretaceous plesiosaurs in some regards.","PeriodicalId":56059,"journal":{"name":"Swiss Journal of Palaeontology","volume":null,"pages":null},"PeriodicalIF":3.0,"publicationDate":"2024-03-15","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"140154737","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Otoliths of the Gobiidae from the Neogene of tropical America","authors":"Werner W. Schwarzhans, Orangel A. Aguilera","doi":"10.1186/s13358-023-00302-5","DOIUrl":"https://doi.org/10.1186/s13358-023-00302-5","url":null,"abstract":"Otoliths are common and diverse in the Neogene of tropical America. Following previous studies of Neogene tropical American otoliths of the lanternfishes (Myctophidae), marine catfishes (Ariidae), croakers (Sciaenidae), and cusk-eels (Ophidiiformes), we describe here the otoliths of the gobies (Gobiidae). The Gobiidae represent the richest marine fish family, with more than 2000 species worldwide and about 250 in America. In the fossil record too they are the species richest family in the Neogene of tropical America. We have investigated otoliths sampled from Ecuador, Pacific and Atlantic Panama, Atlantic Costa Rica, Dominican Republic, Venezuela, and Trinidad, ranging in age from late Early Miocene (late Burdigalian) to late Early Pleistocene (Calabrian). Most of the studied material originates from the collection expeditions of the Panama Paleontology Project (PPP). Our study represents the first comprehensive record of fossil gobies from America, and we recognize 107 species, of which 51 are new to science, 35 are in open nomenclature, and 19 represent species that also live in the region today. Previously, only two fossil otolith-based goby species have been described from the Neogene of tropical America. The dominant gobies in the fossil record of the region are from the Gobiosomatini, particularly of genera living over soft bottoms or in deeper water such as Bollmannia, Microgobius, Antilligobius, and Palatogobius. Another purpose of our study is to provide a first comprehensive account of otoliths of the extant Gobiidae of America, which we consider necessary for an adequate identification and interpretation of the Neogene otoliths. We studied otoliths of 130 extant American gobiid species and figured 106 of them for comparison. We also present a morphological analysis and characterization of the extant otoliths as a basis for the identification of fossil otoliths. Problems that commonly arise with the identification of fossil otoliths and specifically of fossil goby otoliths are addressed and discussed. A comparison of the history of the Gobiidae in tropical America reveals a high percentage of shared species between the Pacific and the Atlantic basins during the Late Miocene (Tortonian and Messinian) from at least 11 to 6 Ma. A recording gap on the Pacific side across the Pliocene allows a comparison again only in the late Early Pleistocene (Calabrian, 1.8 to 0.78 Ma), which shows a complete lack of shared species. These observations support the effective closure of the former Central American Seaway and emersion of the Isthmus of Panama in the intervening time. Groups that today only exist in the East Pacific were also identified in the Miocene and Pliocene of the West Atlantic, and there is also at least one instance of a genus now restricted to the West Atlantic having occurred in the East Pacific as late as the Pleistocene. The evolution of gobies in tropical America and the implications thereof are extensively discussed. Furthermore","PeriodicalId":56059,"journal":{"name":"Swiss Journal of Palaeontology","volume":null,"pages":null},"PeriodicalIF":3.0,"publicationDate":"2024-03-06","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"140044083","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Digital skull anatomy of the Oligocene North American tortoise Stylemys nebrascensis with taxonomic comments on the species and comparisons with extant testudinids of the Gopherus–Manouria clade","authors":"Serjoscha W. Evers, Zahra Al Iawati","doi":"10.1186/s13358-024-00311-y","DOIUrl":"https://doi.org/10.1186/s13358-024-00311-y","url":null,"abstract":"The anatomy of North American tortoises is poorly understood, despite a rich fossil record from the Eocene and younger strata. Stylemys nebrascensis is a particularly noteworthy turtle in this regard, as hundreds of specimens are known from Oligocene deposits, and as this species is one of the earliest fossil turtles to have been described in the scientific literature. Since its initial description based on a shell, many specimens with more complete material have been referred to Stylemys nebrascensis. Here, we review and confirm the referral of an important historic specimen to Stylemys nebrascensis, which includes shell, non-shell postcranial, and skull material. This allows us to document unique skull features of Stylemys nebrascensis (e.g., an unusual ‘poststapedial canal’ that connects the posterior skull surface with the cavum acustico-jugulare) and to refer another well-preserved skull to the species. Based on computed-tomography scanning of these two skulls, we provide a detailed description of the cranial and mandibular osteology of Stylemys nebrascensis. Stylemys nebrascensis has a combination of plesiomorphic skull characteristics (e.g., retention of a medial jugal process) and derived traits shared with extant gopher tortoises (e.g., median premaxillary ridge) that suggest it may be a stem-representative of the gopher tortoise lineage. This supports the hypothesis that extant and fossil tortoises from North America form a geographically restricted clade that split from Asian relatives during the Paleogene.","PeriodicalId":56059,"journal":{"name":"Swiss Journal of Palaeontology","volume":null,"pages":null},"PeriodicalIF":3.0,"publicationDate":"2024-03-05","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"140034083","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Christian Klug, Stephan N. F. Spiekman, Dylan Bastiaans, Beat Scheffold, Torsten M. Scheyer
{"title":"The marine conservation deposits of Monte San Giorgio (Switzerland, Italy): the prototype of Triassic black shale Lagerstätten","authors":"Christian Klug, Stephan N. F. Spiekman, Dylan Bastiaans, Beat Scheffold, Torsten M. Scheyer","doi":"10.1186/s13358-024-00308-7","DOIUrl":"https://doi.org/10.1186/s13358-024-00308-7","url":null,"abstract":"Marine conservation deposits (‘Konservat-Lagerstätten’) are characterized by their mode of fossil preservation, faunal composition and sedimentary facies. Here, we review these characteristics with respect to the famous conservation deposit of the Besano Formation (formerly Grenzbitumenzone; including the Anisian–Ladinian boundary), and the successively younger fossil-bearing units Cava inferiore, Cava superiore, Cassina beds and the Kalkschieferzone of Monte San Giorgio (Switzerland and Italy). We compare these units to a selection of important black shale-type Lagerstätten of the global Phanerozoic plus the Ediacaran in order to detect commonalities in their facies, genesis, and fossil content using principal component and hierarchical cluster analyses. Further, we put the Monte San Giorgio type Fossillagerstätten into the context of other comparable Triassic deposits worldwide based on their fossil content. The results of the principal component and cluster analyses allow a subdivision of the 45 analysed Lagerstätten into four groups, for which we suggest the use of the corresponding pioneering localities: Burgess type for the early Palaeozoic black shales, Monte San Giorgio type for the Triassic black shales, Holzmaden type for the pyrite-rich black shales and Solnhofen type for platy limestones.","PeriodicalId":56059,"journal":{"name":"Swiss Journal of Palaeontology","volume":null,"pages":null},"PeriodicalIF":3.0,"publicationDate":"2024-03-04","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"140034034","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"The first Cyclida from the Triassic of Italy","authors":"Vittorio Pieroni","doi":"10.1186/s13358-024-00306-9","DOIUrl":"https://doi.org/10.1186/s13358-024-00306-9","url":null,"abstract":"A well-preserved carapace of the crustacean Halicyne is here described. The finding comes from the Sostegno Basin (Piedmont, Italy). This is the first occurrence of a well-preserved arthropod from the Middle Triassic San Salvatore Formation of the Biellese area and the first report of a Triassic Cyclida from Italy. Cyclida often occurs in shallow marine environments with rapidly changing salinity conditions. The finding from the Sostegno area suggests a hypersaline paleoenvironment similar to that of Monte San Salvatore (Ticino, Switzerland) upper levels, where the same taxon was previously documented. These fossiliferous localities are shortly compared with that of Rasa di Varese (Lombardy, Italy).","PeriodicalId":56059,"journal":{"name":"Swiss Journal of Palaeontology","volume":null,"pages":null},"PeriodicalIF":3.0,"publicationDate":"2024-02-28","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"140002101","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Bernhard Peyer and his discoveries of Triassic vertebrates in Switzerland","authors":"Hans-Dieter Sues","doi":"10.1186/s13358-024-00310-z","DOIUrl":"https://doi.org/10.1186/s13358-024-00310-z","url":null,"abstract":"Bernhard Peyer (1885–1963) was a prominent Swiss vertebrate palaeontologist and anatomist at the University of Zurich. In 1919, he discovered the occurrence of Middle Triassic (Anisian–Ladinian) ichthyosaurs, sauropterygians, and other reptiles at Monte San Giorgio on the border between Switzerland and Italy. Starting in 1924, Peyer, and later his student and successor Emil Kuhn-Schnyder, worked tirelessly to excavate and collect material from fossil-bearing deposits in this region. The discovery of an articulated skeleton of a long-necked reptile in 1929 allowed Peyer to solve the riddle of unusually elongate bones from the German Muschelkalk, which, he demonstrated, were cervical vertebrae of the extraordinarily long-necked reptile Tanystropheus. As a student in 1919 and later in 1942, Peyer explored a Rhaetian bonebed close to his native town of Schaffhausen. Processing the sediment, he recovered numerous small bones and teeth of vertebrates, among which teeth of diverse stem-mammals are of particular importance.","PeriodicalId":56059,"journal":{"name":"Swiss Journal of Palaeontology","volume":null,"pages":null},"PeriodicalIF":3.0,"publicationDate":"2024-02-28","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"140002108","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"A report on palaeontological excavations and sampling in mudrocks: some guidelines","authors":"Walter Etter","doi":"10.1186/s13358-024-00305-w","DOIUrl":"https://doi.org/10.1186/s13358-024-00305-w","url":null,"abstract":"More than 60% of the world’s sedimentary rocks are mudrocks (Potter et al., 1980; Schieber, 1998; Potter, 2003; the term mudrock is favored here over mudstone because the latter term was used to characterize a limestone texture; Dunham, 1962). From a palaeontological perspective these are, compared to sandstones and limestones, heavily undersampled. The main reason for this is that mudrocks decay in surface exposures to small chips, which develop with sun/heat and rain into an awkward pulp. The decay of the mudstone concomitantly destroys all macrofossils which are not durable. This comprises fossils with an aragonitic or delicate calcitic shell unless they are preserved as pyritic or internal calcitic molds or preserved within calcareous concretions. Therefore, most of the fossils are not recorded in surface exposures. In addition, sedimentologic investigations of mudrocks are hampered because (i) compaction makes sedimentary structures hardly recognizableand (ii) good thin sections of mudrocks are exceedingly difficult to manufacture. For micropalaeontological investigations, mudrocks rich in organic material are especially difficult to process. Standard treatments with boiling water, sodium carbonate solution, or peroxide H2O2 generally fail to dissolve much of the sediment so that the fine fraction (and in mudrocks we usually need the 63μm-fraction) largely consists of clay particle aggregates. Yet there are methods to dissolve these aggregates. Otherwise, picking the microfossils would become extremely laborious. In this paper, some guidelines for successful palaeontological work in mudrocks are outlined. These are based on the author’s personal experience. Examples from Jurassic mudrocks of Switzerland/Europe show that such excavations can be very rewarding.","PeriodicalId":56059,"journal":{"name":"Swiss Journal of Palaeontology","volume":null,"pages":null},"PeriodicalIF":3.0,"publicationDate":"2024-02-15","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"139754236","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Simoniteuthis, a new vampyromorph coleoid with prey in its arms from the Early Jurassic of Luxembourg","authors":"Dirk Fuchs, Robert Weis, Ben Thuy","doi":"10.1186/s13358-024-00303-y","DOIUrl":"https://doi.org/10.1186/s13358-024-00303-y","url":null,"abstract":"The evolutionary history of the cephalopod order Vampyromorpha with its only recent representative, Vampyroteuthis infernalis, the deep-sea vampire squid, is still obscure and a new specimen from the Early Jurassic of Luxembourg, provides new information on the vampyromorph morphology at this period. The new taxon Simoniteuthis michaelyi. gen. n. sp., which is based on a nearly complete gladius with associated head–arm complex, is morphologically intermediate between the families Loligosepiidae and Geopeltidae. Interestingly, the arm crown displays only four arm pairs, although an arm configuration consisting of five arm pairs should be expected in vampyromorph stem lineage representatives. This observation encouraged us to critically review the presumed homology of the filaments of Vampyroteuthis and the lost arm pair in cirrate and incirrate octopods. Moreover, two bony fishes in the mouth region implicated that Simoniteuthis michaelyi n. gen. n. sp. preyed upon them in hostile water depths, a taphonomic phenomenon called distraction sinking. By contrast to modern Vampyroteuthis infernalis, Simoniteuthis michaelyi n. gen. n. sp. roamed and hunted in shallower waters as typical for Mesozoic stem lineage vampyromorphs. According to the current fossil record, a vertical migration into deeper waters (probably associated with a shift in feeding behaviour) occurred at least since the Oligocene.","PeriodicalId":56059,"journal":{"name":"Swiss Journal of Palaeontology","volume":null,"pages":null},"PeriodicalIF":3.0,"publicationDate":"2024-02-14","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"139753752","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Robert Weis, Dominique Delsate, Christian Klug, Thodoris Argyriou, Dirk Fuchs
{"title":"Pachycormid fish fed on octobrachian cephalopods: new evidence from the ‘Schistes bitumineux’ (early Toarcian) of southern Luxembourg","authors":"Robert Weis, Dominique Delsate, Christian Klug, Thodoris Argyriou, Dirk Fuchs","doi":"10.1186/s13358-023-00295-1","DOIUrl":"https://doi.org/10.1186/s13358-023-00295-1","url":null,"abstract":"A re-examination of the early Toarcian fish fossils preserved in public paleontological collections in Luxembourg revealed 70 specimens of large Toarcian pachycormid fish with an excellent three-dimensional preservation within calcareous nodules. Six of them are associated with octobrachian coleoid gladii in their oesophagus or stomach, an association not previously described from Luxembourg. The pachycormids are ascribed to Pachycormus macropterus (Blainville, 1818) and Saurostomus esocinus Agassiz, 1843 while the octobrachian gladii are ascribed to Teudopsis bollensis Voltz, 1836, Teudopsis sp. indet. and Loligosepiidae indet. The position and orientation of the gladii provide direct evidence of these fishes feeding on coleoids and thus a teuthophagous diet, rather than an accidental joint burial. Together with evidence from coeval deposits in Germany, these findings suggest that teuthophagy was a widespread feeding strategy at the base of the clade that contains the suspension-feeding pachycormid giants of the Jurassic–Cretaceous.","PeriodicalId":56059,"journal":{"name":"Swiss Journal of Palaeontology","volume":null,"pages":null},"PeriodicalIF":3.0,"publicationDate":"2024-02-09","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"139753998","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}