Osvaldo A. Conde , Roberto R. Pujana , Gustavo A. Correa , M. Lucía Balarino
{"title":"Barrealoxylon nelsonii gen. et sp. nov., a new Pennsylvanian (Carboniferous) fossil stem from Gondwana: Botanical affinity and paleoclimatic implications","authors":"Osvaldo A. Conde , Roberto R. Pujana , Gustavo A. Correa , M. Lucía Balarino","doi":"10.1016/j.revpalbo.2025.105476","DOIUrl":"10.1016/j.revpalbo.2025.105476","url":null,"abstract":"<div><div>This contribution describes a new gymnospermous fossil stem from western Gondwana. <em>Barrealoxylon nelsonii</em> nov. gen. et sp. was recovered from the Pituil Formation (Early Pennsylvanian), Calingasta-Uspallata Basin in San Juan Province, Argentina. The new species is characterized by a septate pith and mesarch primary xylem. The secondary xylem shows distinct growth ring boundaries, tracheids with uniseriate to triseriate araucarian pits in radial walls, pits in tangential walls, and cross-fields with 1–2, rarely 3, “window-like” (=<!--> <!-->fenestriform) pits. The combination of a septate pith and the primary and secondary xylem characteristics supports its assignment to the Cordaitales, providing further evidence for the presence of this group in southwestern Gondwana. The secondary xylem of the new stem has D-type growth rings, indicative of uniform growth conditions followed by a final stage of reduced vascular cambium activity. This type of growth ring is consistent with the high-humidity climatic conditions that prevailed during the Terminal Glacial Stage in western Gondwana.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"345 ","pages":"Article 105476"},"PeriodicalIF":1.7,"publicationDate":"2026-02-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145571944","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
P.E. Santamarina , V.D. Barreda , M.S. González Estebenet , V. Guler , E.S. Bellosi , F. Novas
{"title":"Palynological analysis of the La Anita and Cerro Fortaleza formations (Campanian), southern margin of the Viedma Lake, Santa Cruz Province, Argentina","authors":"P.E. Santamarina , V.D. Barreda , M.S. González Estebenet , V. Guler , E.S. Bellosi , F. Novas","doi":"10.1016/j.revpalbo.2025.105475","DOIUrl":"10.1016/j.revpalbo.2025.105475","url":null,"abstract":"<div><div>We present the palynological assemblages recovered from the upper part of the La Anita and Cerro Fortaleza formations (Campanian) from SW Patagonia, Argentina. A total of 73 continental species and 11 dinoflagellate cysts species were identified in this study. Continental species comprise 17 bryophytes and lycophytes, 20 ferns, 14 gymnosperms, 21 angiosperms, and 1 of uncertain affinity. The palynofloras are strongly dominated by gymnosperms, with angiosperms and ferns as secondary components. Cluster analysis employed to explore floristic similarities with Campanian–Maastrichtian palynofloras reveals two main groups: one grouping the palynofloras presented here and another grouping Campanian–Maastrichtian palynofloras from southern South America and the Antarctic Peninsula. The occurrence of spores attributable to Salviniales and Anemiaceae, along with pollen grains referable to Symplocaceae, Aquifoliaceae, and Arecaceae suggests a humid and temperate climate. The presence of <em>Odontochitina porifera</em> and <em>Nelsoniella aceras</em> suggests a late Santonian–early Maastrichtian age of deposition. In addition, continental palynological assemblages are consistent with the Campanian age established by radiometric dating. A comparative assessment of angiosperm diversity and abundance within these assemblages and published records indicates that angiosperms had not yet become the dominant floral component in southern Patagonia during the Late Cretaceous, in contrast to coeval floras at similar paleolatitudes in the Northern Hemisphere. However, further high-resolution palynological analyses are needed to refine our understanding of floristic composition, biogeographic patterns, and the time and mode of angiosperm diversification in southern Patagonia during the late Cretaceous.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"345 ","pages":"Article 105475"},"PeriodicalIF":1.7,"publicationDate":"2026-02-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145618568","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Marine Rousseau , Philippe Sorrel , Benjamin Diètre , Morana Čaušević-Bully , Hervé Richard
{"title":"Vegetation changes and human activities since the Bronze Age (late Holocene) in a coastal doline of the Kvarner archipelago (Croatia)","authors":"Marine Rousseau , Philippe Sorrel , Benjamin Diètre , Morana Čaušević-Bully , Hervé Richard","doi":"10.1016/j.revpalbo.2025.105483","DOIUrl":"10.1016/j.revpalbo.2025.105483","url":null,"abstract":"<div><div>We present the first high-resolution palynological and multiproxy reconstruction from a coastal doline in the Kvarner Archipelago (northeastern Adriatic). A continuous sediment core from Osor Lake documents more than three millennia of vegetation change, human impact, and hydrological shifts since the Bronze Age. The record reveals three major phases of landscape transformation: (i) dense deciduous forests (3400–2550 cal BP) with initial signs of clearance; (ii) a progressive landscape opening from 2550 cal BP, marked by expanding grasslands and cultivated fields; and (iii) intensified deforestation and arboriculture after ∼<!--> <!-->800 cal BP. The introduction of <em>Juglans</em> and <em>Castanea sativa</em> between 2200 and 1200 cal BP, together with increasing <em>Olea europaea</em>, reflects evolving Mediterranean agricultural strategies. Charcoal peaks coincide with forest decline, indicating recurrent fire use for land management. This study establishes Osor Lake as a benchmark record for late Holocene land-use history in the northeastern Adriatic and demonstrates how human activities progressively reshaped Mediterranean karst landscapes.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"345 ","pages":"Article 105483"},"PeriodicalIF":1.7,"publicationDate":"2026-02-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145737101","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Ting Wang , Chong Dong , Jianguo Hui , Xueqiong Wei , Gongle Shi
{"title":"Palaeotorreya, a new genus of extinct Taxaceae from the Early Cretaceous of Northeast China","authors":"Ting Wang , Chong Dong , Jianguo Hui , Xueqiong Wei , Gongle Shi","doi":"10.1016/j.revpalbo.2025.105461","DOIUrl":"10.1016/j.revpalbo.2025.105461","url":null,"abstract":"<div><div>Taxaceae (the yew family) are a relict family of conifers, with most species having a highly restricted or fragmentary distribution in the northern hemisphere and New Caledonia. Fossil data indicate that the family had diverged by the earliest Jurassic in southern Sweden, and extant taxaceous genera might had appeared since the Jurassic. In this article, we describe a new genus and species of extinct Taxaceae, <em>Palaeotorreya shenghuii</em> gen. et sp. nov., based on lignified leaves from the Lower Cretaceous Huolinhe Formation in West Ujimqin Banner, eastern Inner Mongolia, Northeast China. The leaves of <em>Palaeotorreya shenghuii</em> are linear to linear-lanceolate and hypostomatic. Epidermal cells of adaxial epidermis and non-stomatal zones of abaxial epidermis are strongly elongated. Two lateral stomatal bands on the abaxial leaf surface are narrow and sunken, each is flanked and concealed by two folding edges bearing dense and prominent papillae. Stomata are longitudinally oriented and arranged in short longitudinal files. Stomata are haplocheilic, stomatal complexes are monocyclic, with a complete and solid stomatal ring surrounding the stomatal pit on the outer cuticle surface. <em>Palaeotorreya shenghuii</em> is most comparable with <em>Torreya</em> within extant conifers, but also bears resemblance in stomatal structure to other genera of extant Taxaceae. The new fossils add to our knowledge of the diversity of the yew family during the Early Cretaceous.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"344 ","pages":"Article 105461"},"PeriodicalIF":1.7,"publicationDate":"2026-01-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145362235","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Elia Ramírez-Arriaga , Aarón Rodríguez-Reyes , Mercedes B. Prámparo , Azucena Canto , Laura Elena Gómez-Lizárraga , G. Iriliana López-Caballero , Alfonso Valiente-Banuet
{"title":"Cactaceae fossil pollen grains recovered from 15.6-Ma evaporites in south-central Mexico","authors":"Elia Ramírez-Arriaga , Aarón Rodríguez-Reyes , Mercedes B. Prámparo , Azucena Canto , Laura Elena Gómez-Lizárraga , G. Iriliana López-Caballero , Alfonso Valiente-Banuet","doi":"10.1016/j.revpalbo.2025.105463","DOIUrl":"10.1016/j.revpalbo.2025.105463","url":null,"abstract":"<div><div>Cacti constitute one of the most iconic and highly diversified groups of plants in the Americas, as Mexico, the Andean region, and eastern Brazil are the main centers of diversity. For many years, its divergence time has been the central question and an enigmatic topic since research has always been hindered by the scarcity of fossil records. The present study documents the palynological diversity of tricolpate and pantocolpate pollen grains with Cactaceae affinity during the Middle Miocene in the Tehuacán-Cuicatlán Valley, an important center of cactus diversity in south-central Mexico. Aridity during this period coincides with the Mid-Miocene Climatic Optimum and is supported by evaporitic rocks from the upper part of the Tehuacán Formation (TF). Tricolpate pollen grains similar to <em>Cephalocereus-Mammillaria</em> cactus genus<em>,</em> were less abundant in the TF, and cluster analysis distinguished three groups. Furthermore, principal component analysis (PCA) showed that the polar area index, the distance between the apices of the colpi, and the equatorial diameter were the characteristics that explained the maximum variance among the tricolpate specimens. In addition, three groups were identified among the pantocolpate specimens with affinities to the genus <em>Pereskia-Opuntia</em>. PCA of the pantocolpate pollen grains revealed that pollen diameter, columella diameter and density, columella length, exine thickness, and spinule density explained the maximum variance among these grains. Miocene pollen grains from Cactaceae were shorter than those of modern species, suggesting that adaptations to the arid climate may have triggered polyploidy and an increase in pollen grain size.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"344 ","pages":"Article 105463"},"PeriodicalIF":1.7,"publicationDate":"2026-01-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145465449","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Searching for Neoproterozoic meteorite impact combusted carbon cenospheres in palynological records","authors":"Morten Smelror","doi":"10.1016/j.revpalbo.2025.105439","DOIUrl":"10.1016/j.revpalbo.2025.105439","url":null,"abstract":"<div><div>Carbon cenospheres comparable to those derived from incomplete combustion of coal or fuel-oil droplets appear to be present in Neoproterozoic post-impact sediments in the Gardnos meteorite crater in central Norway and in possible time-equivalent strata in East Finnmark, northern Norway. The Neoproterozoic cenospheres resemble carbon cenospheres found in K-Pg boundary sediments around the world linked to the Chicxulub meteorite impact on Yucatan and in sediments from the Early Cretaceous Mjølnir impact in the Barents Sea. A likely source of the Late Neoproterozoic carbonized cenospheres is hydrocarbons combusted from organic-rich sediments when the bolide hit the shallow marine impact site of the Gardnos meteorite crater.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"344 ","pages":"Article 105439"},"PeriodicalIF":1.7,"publicationDate":"2026-01-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144902864","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Deep learning of pollen images under low annotation costs: joint optimization of morphological features and training and prediction strategies","authors":"Teng Zhang , Limi Mao","doi":"10.1016/j.revpalbo.2025.105458","DOIUrl":"10.1016/j.revpalbo.2025.105458","url":null,"abstract":"<div><div>Pollen identification is of great importance in the fields of palynology, palaeoecology, botany, medicine and forensic science, but traditional microscopic morphological analysis methods are inefficient and subjective. In this study, we propose an innovative approach based on deep learning to improve the accuracy and efficiency of pollen identification. We constructed a high-quality pollen dataset containing 5521 images of 141 species and a structured attribute table containing 20 standardized morphological features. With an improved ResNet50 architecture, the model utilizes a masking mechanism to combine image features with morphological data, significantly improving classification performance. In addition, we propose a joint training strategy that utilizes both weakly labeled data (unlabeled images + some morphological features) and fully labeled data to alleviate the data scarcity problem. The experimental results show that with the introduction of morphological features, the accuracy of the model significantly improves from 83.00% to at least 89.49% and exhibits stronger generalization ability, effectively reducing overfitting. This study provides a scalable solution for automated pollen identification, addressing key challenges in data utilization and classification accuracy.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"344 ","pages":"Article 105458"},"PeriodicalIF":1.7,"publicationDate":"2026-01-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145362232","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"New data on medullosalean foliage Odontopteris schlotheimii from the uppermost Carboniferous–lowermost Permian of central and eastern Europe","authors":"Zbyněk Šimůnek , Christopher J. Cleal","doi":"10.1016/j.revpalbo.2025.105450","DOIUrl":"10.1016/j.revpalbo.2025.105450","url":null,"abstract":"<div><div>The stomatal structure of the medullosalean frond species <em>Odontopteris schlotheimii</em> is documented for the first time. This supports the view that it is taxonomically distinct from the morphologically similar but rather older frond-species <em>Odontopteris cantabrica.</em> Evidence is also presented that supports the view that the types of <em>Neuropteris pseudoblissii</em> are conspecific with <em>O. schlotheimii</em>.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"344 ","pages":"Article 105450"},"PeriodicalIF":1.7,"publicationDate":"2026-01-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145265929","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"An 11,000-year vegetation and climate record from a subalpine peatland in the Luoxiao Mountains, subtropical Southeast China","authors":"Xin Zhong , Caifeng Cen , Cong Chen , Aiyan Han , Yongjie Tang , Kangyou Huang","doi":"10.1016/j.revpalbo.2025.105442","DOIUrl":"10.1016/j.revpalbo.2025.105442","url":null,"abstract":"<div><div>The evolution of broadleaved forests and their responses to climate changes in subtropical Southeast China remain largely unclear, partly because of the controversies regarding the history of climate (i.e., temperature and monsoonal precipitation) changes during the Holocene. In this study, we integrated pollen and other proxies (including loss on ignition, humification degree, and stable carbon isotope) from a subalpine peatland in the Luoxiao Mountains, to reveal the predominant patterns of vegetation and climate dynamics in mountainous Southeast China. Vegetation at the study site was dominated by <em>Quercus</em>-evergreen and <em>Fagus</em> with relatively flourishing herbs during the early Holocene. A transformation from mixed evergreen–deciduous broadleaved forests to evergreen broadleaved forests occurred between 8200 and 1850 cal. yr BP. The lag of the transformation in response to monsoonal rainfall changes in mountainous regions during the early Holocene can be attributed to thermal limitation. Since 1850 cal. yr BP, broadleaved pollen substantially declined with the expansion of Poaceae and pines, likely reflecting the impact of intensive anthropogenic activities. The expansion of evergreen broadleaved forests from different sites revealed a gradual warming in Southeast China during the Holocene. Additionally, monsoon precipitation in the study area underwent substantial changes, showing a pattern of wetting, sustained humid conditions, subsequent drying, and wetting. Overall, temperature and precipitation during the Holocene were decoupled at the study site, as supported by widely distributed records in mountainous subtropical Southeast China. This may be attributed to the influence of different driving factors on temperature and precipitation.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"344 ","pages":"Article 105442"},"PeriodicalIF":1.7,"publicationDate":"2026-01-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145019472","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Mohammad Firoze Quamar , Nagendra Prasad , Biswajeet Thakur , Maneesha Muraleedharan ET
{"title":"Assessment of herbivore densities using Sporormiella as a palynological proxy: Indian context and the way forward","authors":"Mohammad Firoze Quamar , Nagendra Prasad , Biswajeet Thakur , Maneesha Muraleedharan ET","doi":"10.1016/j.revpalbo.2025.105466","DOIUrl":"10.1016/j.revpalbo.2025.105466","url":null,"abstract":"<div><div>Herbivory plays a crucial role as a long-term driver of ecosystem changes. Understanding the interactions between herbivores and ecosystems is particularly important for shaping conservation policies, especially those focused on re-wilding. Additionally, understanding changes in the abundance of large herbivores over time and assessing the role of key animal species in disturbance regimes, biodiversity dynamics, and ecosystem resilience remains a significant challenge in long-term ecological studies. Spores specific to herbivore dung offer a valuable source of information on past herbivore populations. <em>Sporormiella</em>, a coprophilous fungal spore that grows on herbivore dung, serves as an important proxy for reconstructing the presence and/or abundance of megaherbivores, livestock density, grazing activity (pastoralism), human impact, extinction events, and the effects of husbandry practices on the natural environment over time and across regions. In India, the application of <em>Sporormiella</em> as a proxy for herbivore presence and abundance remains at an early stage and requires further systematic testing. This review article aims to evaluate the potential of <em>Sporormiella</em> in reconstructing herbivore presence and abundance across different temporal and spatial scales in India. The synthesis of <em>Sporormiella</em> records from both modern and the Holocene sediment samples provides insights into past herbivore densities. Additionally, this study discusses various factors influencing the composition of the spores of coprophilous fungi, and the abundance of <em>Sporormiella</em>, potential constraints on their preservation in certain settings, and the limitations of current research. This research review article also outlines broader implications and directions for the future research.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"344 ","pages":"Article 105466"},"PeriodicalIF":1.7,"publicationDate":"2026-01-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145465450","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}