H. Saleh , P. Meyvisch , F. Di Rita , A. Spina , G. Margaritelli , D. Magri , M. Ghilardi , M. Alunni Cardinali , A. Di Michele
{"title":"Evaluating Quercus pollen as a valuable archive of past UV-B levels in the Central Mediterranean: Insights from comparative infrared spectroscopy analyses","authors":"H. Saleh , P. Meyvisch , F. Di Rita , A. Spina , G. Margaritelli , D. Magri , M. Ghilardi , M. Alunni Cardinali , A. Di Michele","doi":"10.1016/j.revpalbo.2025.105427","DOIUrl":"10.1016/j.revpalbo.2025.105427","url":null,"abstract":"<div><div>The relative abundance of ultraviolet-absorbing compounds (UACs) in pollen and spore exine increases with prolonged and higher exposure to ultraviolet B (UV-B) radiation. This relationship has been extensively studied via transmission Fourier-transform infrared microspectroscopy (transmission micro-FTIR), primarily in <em>Lycopodium</em> spores and airborne <em>Pinus</em> pollen. However, traditional transmission micro-FTIR methods are prone to infrared light scattering and interference, resulting in spectral deformations and reduced reproducibility. Additionally, bisaccate pollen, like <em>Pinus</em>, can travel long distances and may not accurately reflect local UV-B levels. This study compares transmission and attenuated total reflection (ATR) micro-FTIR methods to assess their reproducibility and investigates <em>Quercus</em> pollen as a potential local UV-B proxy in the Central Mediterranean. Samples containing fresh (in situ), trapped (in mosses), and fossilized (from Holocene sediments) pollen grains were subjected to various chemical treatments, after which the macromolecular composition of single and clustered grains was characterized. Results show that ATR micro-FTIR yields significantly more reproducible data than transmission micro-FTIR, demonstrating the suitability of this method for systematic chemo-palynological studies. <em>Quercus ilex</em> pollen consistently display UAC-related absorption bands across modern and fossil samples, and treatment with hydrochloric acid, hydrofluoric acid, and sodium hydroxide does not significantly alter these signatures. A comparison with fresh and trapped <em>Q. cerris</em> and <em>Q. pubescens</em> pollen further supports these findings. These results suggest that relative UAC concentrations in modern and fossil <em>Quercus</em> exine likely reflect accumulated UV-B dosage, pinpointing <em>Quercus</em> pollen as a valuable local UV-B proxy for reconstructing past UV-B levels in the Central Mediterranean.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"344 ","pages":"Article 105427"},"PeriodicalIF":1.7,"publicationDate":"2025-08-13","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144908936","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Silvia Ulrich , Carola Purgina , Johannes Martin Bouchal , Christian Geier , Friðgeir Grímsson
{"title":"The use of transmission electron microscopy when investigating fossil angiosperm pollen: A review and suggestions for future applications","authors":"Silvia Ulrich , Carola Purgina , Johannes Martin Bouchal , Christian Geier , Friðgeir Grímsson","doi":"10.1016/j.revpalbo.2025.105423","DOIUrl":"10.1016/j.revpalbo.2025.105423","url":null,"abstract":"<div><div>It has been more than 65 years since transmission electron microscopy (TEM) was first used to study fossil angiosperm pollen. Since then, significant progress has been made in sample preparation protocols and TEM equipment, particularly in contrasting methods used to reveal detailed ultrastructural differences in pollen walls, as well as sensitive, high-resolution cameras needed for documentation. Early TEM studies on fossil angiosperm pollen focused on determining whether ultrastructure could be used to assign palynomorphs to spore-producing plants, gymnosperms, or angiosperms. Later research became more taxonomically focused, comparing fossil pollen walls to those of extant angiosperms. Most TEM research on fossil angiosperm pollen has until now been conducted in the USA, Western/Central Europe, and Russia, with limited contributions from other parts of the scientific community. There is a clear relation between the sample origin (country, locality) and author group, and between the type of pollen specimens (dispersed <em>versus in situ</em>) and the taxonomic resolution. <em>In situ</em> pollen from fossil flowers is usually assigned to families and genera, while most dispersed pollen is noted as <em>Incertae sedis</em>. Despite the obvious advantages of TEM analyses for determining the taxonomic placement and phylogenetic relations of fossil pollen, little effort has been put into this subject over the last decades. To increase the understanding of angiosperm evolution, TEM investigations of fossil pollen need to be routinely applied. New generations of palynologists need to pick up on this method, and an effort to investigate Cenozoic angiosperm pollen from a taxonomic point of view is overdue.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"344 ","pages":"Article 105423"},"PeriodicalIF":1.7,"publicationDate":"2025-08-06","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144867231","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Iván R. Barreiro , José B. Diez , Anna Fijałkowska-Mader , Elke Schneebeli-Hermann , Amalia Spina , Evelyn Kustatscher
{"title":"Reassessment of the chronostratigraphic and paleogeographic distribution of the Permian prepollen genus Nuskoisporites","authors":"Iván R. Barreiro , José B. Diez , Anna Fijałkowska-Mader , Elke Schneebeli-Hermann , Amalia Spina , Evelyn Kustatscher","doi":"10.1016/j.revpalbo.2025.105425","DOIUrl":"10.1016/j.revpalbo.2025.105425","url":null,"abstract":"<div><div>The genus <em>Nuskoisporites</em> is a common component of Permian palynofloras, with biostratigraphic and paleogeographic importance. Despite the existence of previous revisions of its occurrence, there are still uncertainties concerning its complete stratigraphic range and paleogeographic distribution. This study presents a comprehensive review of historical records, complemented by new morphometric data derived from 446 <em>Nuskoisporites</em> specimens representing ten geographically distinct regions. Three morphometric variables were measured and analyzed using Principal Component Analysis (PCA): total size (diameter), the ratio of total size to central body diameter (P:CB), and the ratio of central body diameter to average laesurae length (CB:L). Two morphotypes can be distinguished based on those characters, each exhibiting different stratigraphic ranges. The older morphotype spans the Pennsylvanian to lower Guadalupian, while the younger form is confined to the upper Cisuralian through Lopingian. Current evidence supports the extinction of <em>Nuskoisporites</em> shortly before or at the Permian–Triassic boundary, thereby reflecting a genuine ecological turnover rather than reworking or preservation bias. Paleogeographic and paleoecological analyses indicate that <em>Nuskoisporites</em> initially had a broad distribution across multiple phytoprovinces and latitudes. However, a marked reduction of its geographic distribution is observed during the Lopingian, restricting the pollen to low-latitude regions of the western Tethys margin, coinciding with intensified global aridification and environmental stress. This integrated approach of <em>Nuskoisporites</em> provides a refined stratigraphic range for the genus and new insights into its ecological dynamics during the end of the Carboniferous and Permian.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"343 ","pages":"Article 105425"},"PeriodicalIF":1.7,"publicationDate":"2025-08-05","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144828211","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
María de las Mercedes Sosa , Mariela Nuñez-Florentín , María Betiana Angulo , Juan Manuel Coronel , Cristina Salgado Laurenti
{"title":"Morphological variation and evolutionary trends in pollen of Bacopa (Gratioleae-Plantaginaceae)","authors":"María de las Mercedes Sosa , Mariela Nuñez-Florentín , María Betiana Angulo , Juan Manuel Coronel , Cristina Salgado Laurenti","doi":"10.1016/j.revpalbo.2025.105422","DOIUrl":"10.1016/j.revpalbo.2025.105422","url":null,"abstract":"<div><div><em>Bacopa</em> (Gratioleae-Plantaginaceae) is a plant genus with great morphological variability and little previous information on its palynology. Thus, the aim of the present study is to describe palynological characters of potential taxonomic significance for the delimitation of <em>Bacopa</em> species and to discuss their systematic implications. In addition, the evolution and systematic utility of palynological characters in the genus were also analyzed. The pollen grains were acetolyzed, observed and analyzed under light and scanning electron microscopy, measured and photographed. Analyses based on qualitative and quantitative data were performed based on descriptive and multivariate statistics. Also, the ancestral reconstruction of palynological characters was carried out by stochastic character mapping, based on previous phylogenies. The pollen morphology of 22 taxa was described for the first time. The pollen morphology of <em>Bacopa</em> contributes to taxonomic studies of the genus because it allows the formation of groups that coincide with previously proposed sections. The compound opening (tricolporate) was considered as an ancestral character state, as well as the tectate exine, small size and prolate-spheroidal shape. <em>Bacopa</em> is confirmed as an eurypalynous genus, the number and type of apertures allowed us to differentiate three pollen types that coincide with the subclades determined by molecular phylogeny.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"343 ","pages":"Article 105422"},"PeriodicalIF":1.7,"publicationDate":"2025-07-28","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144723757","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
N. Hayrapetyan , E. Hakobyan , M. Melkonian , A. Mamyan , E. Kvavadze , G. Zhamakochyan , A.A. Bruch , I. Gabrielyan
{"title":"First record of fossil algal diversity in Lake Sevan, Armenia: Illuminating ecological dynamics and environmental parameters","authors":"N. Hayrapetyan , E. Hakobyan , M. Melkonian , A. Mamyan , E. Kvavadze , G. Zhamakochyan , A.A. Bruch , I. Gabrielyan","doi":"10.1016/j.revpalbo.2025.105420","DOIUrl":"10.1016/j.revpalbo.2025.105420","url":null,"abstract":"<div><div>Algae serve as valuable proxies for paleoenvironmental reconstructions in sedimentary environments. Fossil algae from Middle to Late Holocene sediments, alongside pollen data, enrich our understanding of environmental changes, particularly evident in the Tsovinar-1 section of Lake Sevan, Armenia.</div><div><em>Botryococcus</em> and <em>Pediastrum/Pseudopediastrum</em> are sensitive indicators of hydrological changes and reflect fluctuations in lake level. The comparatively high abundance of <em>Pediastrum/Pseudopediastrum</em> and <em>Botryococcus</em> from 6000 BC to 4400 ca. BC, when water levels were elevated<em>,</em> together with the occurrence of the genera <em>Gomphonema</em>, <em>Fragilaria</em>, <em>Cymbella</em>, and <em>Epithemia</em> during high lake level phases, underscores their utility as pelagic indicators. From 4400 to 2600 ca. BC, no aquatic vegetation is recorded at Tsovinar-1 under relatively more arid conditions. Later, from 2600 to 1400 ca. BC the lake level rose again and eutrophic conditions in some shallow areas along the coastal zone developed, indicated by high abundances of the nitrogen-fixing cyanobacterium <em>Gloeotrichia</em>. Later, after 600 ca. BC, further variations in water level occurred and are mirrored in algal abundances. Changes of pH followed those trends of water level and are mainly reflected in the occurrences of Zygnematophyceae, particularly with high numbers of <em>Spirogyra</em> and <em>Mougeotia</em>.</div><div>The results suggest that algal species hold a valuable potential for paleoecological reconstructions, calling for further studies across diverse environmental gradients. Notably, the first documentation of <em>Gloeotrichia</em> in the fossil record of the Caucasus provides a new proxy for the analysis of environmental changes in lakes and pools of the Armenian Highland.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"343 ","pages":"Article 105420"},"PeriodicalIF":1.7,"publicationDate":"2025-07-23","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144748659","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Andrey G. Fedyaevskiy , Alexey V. Vergunov , Sofia S. Shadrina
{"title":"Palaeoenvironmental response to the Early Cretaceous volcanic ash fall: Evidence from palynology and palynofacies of the tonstein-bearing coal seam of Kharanor Deposit, Transbaikalia, Russia","authors":"Andrey G. Fedyaevskiy , Alexey V. Vergunov , Sofia S. Shadrina","doi":"10.1016/j.revpalbo.2025.105421","DOIUrl":"10.1016/j.revpalbo.2025.105421","url":null,"abstract":"<div><div>Volcanic ash falls, being one of the consequences of eruptive palaeoevents, are reflected in the geological record of coal-bearing sediments as tonsteins, thin clayey interlayers. Information on the palaeontological characteristics of tonsteins and assessments of the influence of ash on the palaeoenvironment are still fragmentary. This paper presents new palynological and palynofacies data obtained for the tonstein-bearing Kharanor coal deposit located in Transbaikalia, Russia. The spore and pollen assemblages indicate the early Albian (Cretaceous) age of the sediments. The tonstein palynomorph composition differs markedly from the host coal one, which is expressed in an increased number of fern spores and a decreased content of gymnosperm pollen. The palynofacies composition of the Kharanor tonstein is characterised by a high content of phytoclasts of the cutinite subgroup and an increased content of palynomorphs and amorphous organic matter, whereas the lower and upper coals are dominated by vitrinite and inertinite phytoclasts. The results indicate peat-forming conditions that existed in a swampy area where a predominantly gymnosperm-fern forest grew in a wet and warm-temperate climate. At one point the area was covered by a relatively thick layer of volcanic ash, resulting in the burial of the lower forest layers, possibly during their spore production period. Rapid rates of sedimentation and subsequent compaction resulted in the conservation of organic matter, including unstable soft plant tissues. However, the impact of the volcanic event on vegetation and palaeoenvironment had a noticeable but short-term effect. No fundamental changes in flora or sedimentation conditions followed in the long term.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"343 ","pages":"Article 105421"},"PeriodicalIF":1.7,"publicationDate":"2025-07-23","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144722832","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"World list of Quaternary pollen and spore atlases, an update","authors":"Henry Hooghiemstra , Suzette G.A. Flantua","doi":"10.1016/j.revpalbo.2025.105404","DOIUrl":"10.1016/j.revpalbo.2025.105404","url":null,"abstract":"<div><div>In the past decades, numerous studies have focused on the morphological identification of fossil pollen grains and fern spores. This paper provides an updated compilation of regional Quaternary pollen and spore floras, categorized per continent. This list includes books, standalone papers, series under a common title, textbooks, and open access electronic pollen reference collections available at websites. Regional pollen floras cover diverse geographical scales, from urban areas and national parks to mountain areas and broader phytogeographical units. This update builds on the ‘World list of Quaternary pollen and spore atlases’ published in 1998, incorporating over 300 new references from 1998 to 2025, bringing the total number of references to almost 800. The quality of photographs varies much, pollen identification keys are not always provided, and some publications remain difficult to access. A thorough understanding of available printed literature and online pollen reference collections is essential for improving the accuracy of fossil pollen and spore identification.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"343 ","pages":"Article 105404"},"PeriodicalIF":1.7,"publicationDate":"2025-07-18","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144766807","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Ao Yan , Jiangyong Wang , Zhankui Bai , Xudong Liu , Qingcao Tian
{"title":"Paludification-driven peatlands development and vegetation variations in Alataw Mountains, Northwest China since the Late Holocene: Insights from multiple indicators","authors":"Ao Yan , Jiangyong Wang , Zhankui Bai , Xudong Liu , Qingcao Tian","doi":"10.1016/j.revpalbo.2025.105419","DOIUrl":"10.1016/j.revpalbo.2025.105419","url":null,"abstract":"<div><div>Exploring the impacts of environmental changes on the peatlands evolution and vegetation succession can provide valuable insights for studying the development of wetland ecosystems, especially in arid Northwest China. This study conducted analyses of phytoliths, grain size, organic matter, and humification degree in the ARXT peatland within the Alataw Mountains, Xinjiang, and further investigated the formation, development, and vegetation dynamics of the peatland over the past 3100 years by integrating vegetation Rate-of-change (RoCs) analysis and structural equation modeling. The results indicate that the period from 3100 to 2600 cal yr BP was characterized by a shallow lacustrine environment, followed by the gradual onset of paludification and incipient peat accumulation. Between 2600 and 1000 cal yr BP, the system transitioned to a swamp environment, during which peat underwent stable development. Since 1000 cal yr BP, the peatland has been subjected to a generally cold and dry environment, leading to accelerated decomposition of organic matter and unstable peat accumulation. With the onset of lake paludification, herbaceous taxa dominated by Poaceae and Cyperaceae underwent rapid proliferation. Vegetation RoCs increased markedly during the lake paludification phase (3100–2362 cal yr BP), stabilized throughout the peat accumulation period (2362–720 cal yr BP), and rose again after 720 cal yr BP, concomitant with environmental cooling and declining peatland water levels. Driven by the joint influence of temperature and wetland hydrology, paludification emerged as the primary driver of ARXT peatland development and vegetation dynamics. Human pasture activities exerted no significant influence on vegetation changes in ARXT peatland.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"343 ","pages":"Article 105419"},"PeriodicalIF":1.7,"publicationDate":"2025-07-18","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144678956","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Deciphering interfungal relationships in the 410-million-yr-old Rhynie chert: Glomites bacatus nov. sp. (sporocarp-forming Glomeromycota) and associated microfungi","authors":"Michael Krings","doi":"10.1016/j.revpalbo.2025.105418","DOIUrl":"10.1016/j.revpalbo.2025.105418","url":null,"abstract":"<div><div>The fossil record of the Glomeromycota (arbuscular mycorrhizal fungi) from the Lower Devonian Rhynie chert of Scotland includes a wide variety of spores, most of which formed singly in the axes of land plants; only a few have been reported to occur as sporocarps. The Rhynie chert also provides ample evidence that these spores were attractive as a habitat and nutritional basis for other fungi. Here, I describe a new glomeromycotan sporocarp from the Rhynie chert, <em>Glomites bacatus</em> nov. sp., which consists of more than 100 glomoid spores and is completely enveloped by a peridium of tightly interlacing and interlocking hyphae with vesicle-like swellings. Both the peridium and the spores of the specimens at hand are colonized by other fungi, including chytrid-like thalli comparable to <em>Rhizophydites bicornis</em>. Other remains associated with the sporocarps could belong to fungus-like Oomycetes. These include stalked spherules reminiscent of chlamydospores, flask-shaped structures resembling sporangia, and a putative intercalary oogonium containing two unequal-sized oospores and subtended by a hypogynous antheridium with a fertilization tube protruding into one of the oospores. This discovery expands our knowledge of sporocarpic Glomeromycota in the Rhynie ecosystem and provides further evidence for the function of these fungi as hosts for other microbial life.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"343 ","pages":"Article 105418"},"PeriodicalIF":1.7,"publicationDate":"2025-07-17","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144670854","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Henk Brinkhuis , Chiara Fioroni , Mustafa Yücel Kaya
{"title":"There and back again; on dinoflagellate cyst index events of the Eocene - Oligocene Transition in the (Para)Tethyan Realm","authors":"Henk Brinkhuis , Chiara Fioroni , Mustafa Yücel Kaya","doi":"10.1016/j.revpalbo.2025.105414","DOIUrl":"10.1016/j.revpalbo.2025.105414","url":null,"abstract":"<div><div>A recent biochronostratigraphic (coccolithophorids, dinoflagellate cysts) and paleoenvironmental analysis of the hemipelagic deposits of the İhsaniye Formation, exposed along the cliffs in the Karaburun area (Black Sea coast, NW Turkey) provided new insights into the paleoceanographic and paleoclimatic evolution of the central (Para-)Tethyan region across the Eocene–Oligocene Transition (EOT). Among others, the study identified the Earliest Oligocene Stable Isotope Step (EOIS) marking the inception of Antarctica's first continental-scale ice sheets since the mid-Permian and coinciding with a major eustatic lowering, followed by the Early Oligocene Glacial Maximum (EOGM) period with its peak δ<sup>18</sup>O values. The study showed apparent (quasi) continuity of the EOT succession at Karaburun, a notion that is not a 100% obvious from the organic walled dinoflagellate cyst (or dinocyst) record. This is mainly because the iconic Eocene – earliest Oligocene taxon <em>Areosphaeridium diktyoplokum</em>, in coeval sections in the region quite abundant near the end of the Eocene, is virtually absent. Here, we focus on obtaining a more detailed picture of correlative secondary dinocyst and other EOT bioevents allowing an even more robust chronostratigraphic assessment of the succession, including correlation to the Italian type sections. Particularly the ranges of the (herein taxonomically revised) species <em>Explodinium priabonensis</em> gen. and comb. nov. and the new species <em>Glaphyrocysta peterbijlii</em> sp. nov. may be regarded as additional criteria to correlate EOT strata within the (Para)Tethyan realm. Combined evidence now suggests that the Karaburun section may not be as complete as previously assumed and that a small part of the succession correlative to the base of the EOGM, representing a portion of the Adi dinocyst Zone may be missing. This aspect does not affect the overall outcome and significance of the studies at Karaburun. In fact, a brief hiatus may well be driven by the major EOT sea level lowering.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"343 ","pages":"Article 105414"},"PeriodicalIF":1.7,"publicationDate":"2025-07-16","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144662710","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}