Review of Palaeobotany and Palynology最新文献

{"title":"Late Aptian spore-producing plant records in the South Atlantic: Distribution, botanical affinities, ecological and climatic implications","authors":"Marcelo de A. Carvalho,&nbsp;Michelle C.S. Giannerini,&nbsp;Gabriel C. Correia,&nbsp;Cecília C. Lana,&nbsp;Natália de P. Sá,&nbsp;Gustavo Santiago,&nbsp;Viviane S.F. Trindade,&nbsp;Maria Eduarda C. Coelho","doi":"10.1016/j.revpalbo.2026.105531","DOIUrl":"10.1016/j.revpalbo.2026.105531","url":null,"abstract":"<div><div>The late Aptian record of spore-producing plants along the South Atlantic margin provides a robust proxy for reconstructing paleogeographic, ecological, and climatic conditions during the early stages of the South Atlantic opening. A total of 567 samples from eight Brazilian sedimentary basins (Bragança-Viseu, São Luís, Parnaíba, Ceará, Potiguar, Sergipe, Araripe, and Espírito Santo) were analyzed for spore-producing plant families. Eighteen families were identified, including bryophytes (e.g., liverworts, Anthocerotaceae), lycophytes (e.g., Lycopodiaceae, Selaginellaceae), and ferns (e.g., Anemiaceae, Cyatheaceae, Dicksoniaceae, Marsileaceae, Schizaeaceae). Quantitative analyses reveal that Anemiaceae is the most abundant and geographically widespread family, representing over 40% of the total spore-producing plant count. Its dominance, coupled with high Consistency Index (CI) values, indicates strong ecological tolerance and stability under fluctuating climatic regimes. The Pearson correlation matrix shows positive associations among major fern and lycophyte families and negative correlations with xerophytic taxa, reinforcing their link to humid or mesic paleoenvironments. Variations in the ratio of spore-producing plants to xerophytes (SPP/X) delineate three main paleoclimatic phases: pre-evaporitic, evaporitic, and post-evaporitic, each marked by alternating arid and humid intervals. Integration with previous studies suggests that the Intertropical Convergence Zone (ITCZ) and the early configuration of the South Atlantic Ocean played key roles in modulating regional humidity. The observed floral composition highlights the resilience and adaptability of spore-producing plants, particularly Anemiaceae, under transitional climatic regimes of the late Aptian.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"348 ","pages":"Article 105531"},"PeriodicalIF":1.7,"publicationDate":"2026-05-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146191014","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"A seed of Allicospermum angrenicum Nosova from the Middle Jurassic of Uzbekistan with a trapped pollen grain","authors":"Natalya Nosova ,&nbsp;Natalia Zavialova","doi":"10.1016/j.revpalbo.2026.105513","DOIUrl":"10.1016/j.revpalbo.2026.105513","url":null,"abstract":"<div><div>We present new data on the anatomy and the first data on the ultrastructure of all cuticles and the megaspore membrane of a seed of <em>Allicospermum angrenicum</em> from the Middle Jurassic Angren Formation of the Angren locality in Uzbekistan. A transmission electron microscope study reveals the outer and inner cuticles of the integument, the cuticle of the nucellus, and the megaspore membrane. The cuticular membrane of the outer integument is composed of a cuticle homogeneous proper and a cuticular fibrillar layer. The megaspore membrane is made up of a patterned layer and a homogeneous foot layer. The patterned layer is composed of crowded bacula arranged vertically or rarely obliquely to the foot layer that is why some of them look as rounded or oval bodies on transverse or oblique sections. Several homogeneous orbicule-like bodies were observed in the upper part of the megaspore membrane. A pollen grain was found entrapped in the seed. The monosulcate pollen has an ectexine with a solid prominent tectum, a prominent infratectum that shows alternation of alveoli with rounded outlines and transversely elongated alveoli, and a thin insignificant foot layer, and a more electron dense endexine. The information on the seed anatomy and ultrastructure does not exclude both cycadalean and ginkgoalean affinities of the find, whereas the pollen characters definitely point to the cycadalean affinity.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"348 ","pages":"Article 105513"},"PeriodicalIF":1.7,"publicationDate":"2026-05-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146191089","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"The pulse of ancient fires: Reconstructing Toarcian wildfire regimes from fossil charcoals in northwest China","authors":"Yuling Na ,&nbsp;Xiao Shi ,&nbsp;Chi Tian ,&nbsp;Wenpei Pei ,&nbsp;Qianqian Song ,&nbsp;Shaoting Tian ,&nbsp;Bing Wang ,&nbsp;Yan Liu ,&nbsp;Leon Nahuel Torres ,&nbsp;Jianxin Yu","doi":"10.1016/j.revpalbo.2026.105519","DOIUrl":"10.1016/j.revpalbo.2026.105519","url":null,"abstract":"<div><div>Understanding the occurrence and ecological impacts of paleo-wildfires is essential for reconstructing paleoclimate conditions and ecosystem evolution. This study focuses on fossil charcoals from the Sangonghe Formation in the HD1 well of the Hoxtolgay Basin, northern Xinjiang, China. According to the Scanning Electron Microscope (SEM) microscopic characteristics, the charcoal assemblage from the Sangonghe Formation comprises <em>Agathoxylon</em>, <em>Protocupressinoxylon</em>, <em>Protophyllocladoxylon</em>, <em>Protosciadopityoxylon</em>, and gymnosperms incertae sedis. Macro-charcoals are primarily in situ or near in situ, indicating frequent low-intensity local surface fires with temperatures ranging from 282.91 °C to 591.37 °C, mostly concentrated between 300 and 500 °C. The abundance of charcoals and paleo-wildfire temperatures show significant vertical variations, with more active, higher-temperature wildfire activities, and more obvious fire temperature fluctuations in the 523.8–673.6 m interval, possibly related to global surge in wildfires and climatic aridification during the initiation of the Toarcian Oceanic Anoxic Event (T-OAE). The charcoal records from the Sangonghe Formation constrain the atmospheric oxygen concentration to no less than 15% during the late Early Jurassic. Based on the paleo-wildfires reflected by charcoals and palynological characteristics, the climate in the Hoxtolgay Basin during the late Early Jurassic was predominantly warm and humid, probably with intermittent aridity. The sedimentary process of the Sangonghe Formation may have experienced a transition from arid to humid conditions. The present study reports the first systematic taxonomic analysis of the late Early Jurassic charcoals in northern Xinjiang, providing crucial evidence for reconstructing paleo-wildfire dynamics and exploring the coupling mechanisms of wildfires, climate, and ecosystems in geological history.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"348 ","pages":"Article 105519"},"PeriodicalIF":1.7,"publicationDate":"2026-05-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146081611","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"Pollen morphology and ultrastructure of extant Castanoideae (Fagaceae): 1. Castanopsis.","authors":"Natalia Naryshkina ,&nbsp;Maria Tekleva","doi":"10.1016/j.revpalbo.2026.105532","DOIUrl":"10.1016/j.revpalbo.2026.105532","url":null,"abstract":"<div><div>The pollen morphology and ultrastructure of 15 species of <em>Castanopsis</em> (Fagaceae) was studied using transmitted light and electron microscopes. Pollen grains are small, prolate, tricolporate, rugulate. The ectexine consists of a relatively thick tectum, columellate infratectum and distinct foot layer. At the apertural region, the ect- and endexine form specific structures appearing like rollers flanking the apertural region. The principal pollen characters for distinguishing <em>Castanopsis</em> species or group of species are rugula length and width and rugula distribution on the exine surface. Four groups of <em>Castanopsis</em> species were established based on the exine sculpturing. Additional pollen characters for distinguishing are thickness of the ectexine layers and their ratio, and columella width. Obtained information allows making identification of fossil Castanoideae pollen more accurate. Our data partly correspond to the existing systematics of the genus; the systematics of the genus should be improved and palynological data are an important part to be included.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"348 ","pages":"Article 105532"},"PeriodicalIF":1.7,"publicationDate":"2026-05-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146191013","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"Evidence for Early Middle Jurassic wildfires: New macro-charcoal records from the Xishanyao Formation, Junggar Basin, Northern Xinjiang, China","authors":"Bing Yang ,&nbsp;Yuling Na ,&nbsp;Xiao Shi ,&nbsp;Yi Wei ,&nbsp;Xinzhi Zhang ,&nbsp;Bing Wang ,&nbsp;Siyuan Sun ,&nbsp;Weitong Li","doi":"10.1016/j.revpalbo.2026.105518","DOIUrl":"10.1016/j.revpalbo.2026.105518","url":null,"abstract":"<div><div>Wildfires, as an integral part of the Earth system, profoundly influence climate, vegetation, and biogeochemical cycles, shaping global ecological and climatic dynamics. In this study, we systematically analyzed charcoal fossils from the Middle Jurassic Xishanyao Formation in the Junggar Basin using taxonomic classification and reflectance measurements. Our analysis suggests frequent, low-temperature (318–493 °C) and low-intensity surface fires, with the preserved charcoal indicating contributions from <em>Protophyllocaldoxylon</em>-like gymnosperm wood as well as other taxonomically indeterminate gymnosperm fragments. The presence of abundant, large macro-charcoal fragments provides high-fidelity evidence of local wildfire activity, indicating a warm and humid subtropical climate conducive to biomass accumulation that was probably punctuated by intermittent dry intervals. High atmospheric oxygen levels (&gt;<!--> <!-->15%, potentially up to 28.3%) and lightning strikes are considered the primary drivers that facilitated combustion. This research provides important new evidence from the Xishanyao Formation to further refine the reconstruction of paleo-climate and wildfire activity during the Middle Jurassic and offers key insights into the terrestrial ecosystem evolution of Northwest China during this period.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"348 ","pages":"Article 105518"},"PeriodicalIF":1.7,"publicationDate":"2026-05-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146081610","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"A contribution to the palynology and its taxonomic significance in Acanthophyllum (Caryophyllaceae) from Iran","authors":"Mehdi Dehghani ,&nbsp;Alireza Dolatyari","doi":"10.1016/j.revpalbo.2026.105520","DOIUrl":"10.1016/j.revpalbo.2026.105520","url":null,"abstract":"<div><div>Pollen morphology of 76 populations representing 26 species and five sections of <em>Acanthophyllum</em> was investigated using both light and electron microscopy to assess the largely unexplored potential of pollen traits for infrageneric classification within the genus. Extracted from herbarium specimens, the pollen grains were acetolyzed for light microscopy (LM) analysis, while intact grains were used for scanning electron microscopy (SEM). A range of pollen traits was investigated, including pollen and pore diameters, exine thickness, interporal distance (chord), pore diameter/pollen diameter ratio as well as pore number and density. In addition, SEM observations focused on spinule morphology, measuring spinule length and width, as well as the density of spinules and perforations on the exine surface. Pollen grains of <em>Acanthophyllum</em> species are monads, spheroidal to subspheroidal, tectate, pantopolyporate. Both tectum and operculum are ornamented with spinules. Pollen size is medium, ranging from 24.29 to 38.07 μm in diameter, with the average number of pores varying between 31 and 51 among the examined species. While pollen characters showed limited utility for distinguishing species, they provided strong evidence for sectional delimitation, albeit with some overlaps and outliers. Overall, pollen traits were conserved at the generic level but showed sectional differences, with the observed variation aligning with phylogenetic relationships within the genus. Pollen evolution in <em>Acanthophyllum</em> reflects the pressures of polyploidy, morphological plasticity, and adaptation to arid environments, exhibiting general homogeneity but variation in size, aperture number, and spinule density.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"348 ","pages":"Article 105520"},"PeriodicalIF":1.7,"publicationDate":"2026-05-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146191090","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"Re-investigation of Pterophyllum crassinervum (Bennettitales) from the Rhaetian of South China and its palaeoecological implications","authors":"Yuanyuan Xu ,&nbsp;Ning Lu ,&nbsp;Liqin Li ,&nbsp;Yongdong Wang","doi":"10.1016/j.revpalbo.2026.105516","DOIUrl":"10.1016/j.revpalbo.2026.105516","url":null,"abstract":"<div><div><em>Pterophyllum</em> Brongniart 1825 is one of the most abundant morphogenus of bennettitalean foliage with at least 157 species documented in China. Among the numerous <em>Pterophyllum</em> species, <em>P. crassinervum</em> Huang et Chow was originally established solely based on insufficiently macromorphological features with limited fossil records. Here, we re-investigate this species for detailed descriptions and interspecific comparisons, and firstly show its epidermal structure based on new fossil collections from the Rhaetian of Upper Triassic strata in the northeastern Sichuan Basin, South China. The thoroughly re-examination of <em>Pterophyllum crassinervum</em> confirms its taxonomic validity and define the standard criteria for distinctive segregation of this species from its analogues. A neotype and an emended specific diagnosis with supplementary microstructures of <em>Pterophyllum crassinervum</em> are formally proposed. Based on a comprehensive interspecific comparison of <em>Pterophyllum</em> species, we confirm that the occurrence of mucronate teeth on pinnae apices is a rare character among this plant group. Considering the predominant position of Bennettitales (especially <em>Pterophyllum</em>) and active herbivories (especially marginal/apical feeding) during the latest Triassic in northeastern Sichuan Basin, the appearance of mucronate teeth may represents the plant-insect co-evolutionary results.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"348 ","pages":"Article 105516"},"PeriodicalIF":1.7,"publicationDate":"2026-05-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146071136","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"Cuticular evidence and taxonomic reassessment of Miocene Albizia fossils in southwest China: Implications for the biogeography of A. julibrissin","authors":"Ping Lu,&nbsp;Jin-Bo Gao,&nbsp;De-Liang Li,&nbsp;Xiao-Qing Liang","doi":"10.1016/j.revpalbo.2026.105517","DOIUrl":"10.1016/j.revpalbo.2026.105517","url":null,"abstract":"<div><div><em>Albizia julibrissin</em>, native to Asia, has been widely introduced to and cultivated in both America and Europe. Well-preserved fossil leaflets recovered from late Miocene deposits in Huaning, southwest China, exhibit distinctive characteristics consistent with those of extant <em>A. julibrissin.</em> These include: (1) falcate or oblong leaflets with the midvein positioned adjacent to the upper margin, (2) distinctive trichome bases on the lower epidermis, (3) straight to moderately undulate anticlinal cell walls, and (4) anomocytic stomatal complexes. However, because of the smaller size of the epidermal cells observed in the fossil leaflets, these specimens have been reclassified as <em>Albizia</em> cf. <em>julibrissin</em> Durazz. These fossils show that ancestors of <em>A. julibrissin</em> were already present in southwest China during or before the warm late Miocene. During the late Neogene, <em>A. julibrissin</em> spread widely despite prevailing global cooling trends. In contrast, contemporary climate projections suggest that a warming of 1.5 °C by 2100 could lead to a substantial contraction of the suitable range for this warm-adapted Miocene species in the coming decades. This reduction is expected to occur despite efforts in artificial cultivation and human-assisted dispersal.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"348 ","pages":"Article 105517"},"PeriodicalIF":1.7,"publicationDate":"2026-05-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146081609","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"A new Late Pleistocene Trapa (Lythraceae) species from the Nepal Himalaya and its implications for biogeography and paleoenvironment","authors":"Dhan Bahadur Khatri ,&nbsp;Weilin Zhang ,&nbsp;Maodu Yan ,&nbsp;Cuirong Yu ,&nbsp;Xiaomin Fang ,&nbsp;Purushottam Adhikari ,&nbsp;Gaurav Srivastava ,&nbsp;Fuli Wu ,&nbsp;Jinbo Zan ,&nbsp;Khum Narayan Paudayal","doi":"10.1016/j.revpalbo.2026.105521","DOIUrl":"10.1016/j.revpalbo.2026.105521","url":null,"abstract":"<div><div>The genus <em>Trapa</em> L. (Lythraceae) encompasses annual, free-floating plants, predominantly distributed across tropical to warm temperate regions. Despite its extensive modern biogeographic range, the Himalayan fossil record of <em>Trapa</em> remains exceedingly sparse, with previously documented occurrences restricted to the Pliocene-Pleistocene deposits in the western Himalayas. Herein, we describe a newly discovered fossil fruit of <em>Trapa</em> from the Late Pleistocene of the northeastern Kathmandu Basin, Nepal, designated <em>Trapa gokarnansis</em> D.B. Khatri sp. nov. The specimen displays diagnostic morphological characteristics, including a woody pericarp, obovate shape, wider upper two horns, a conspicuously swollen and broadly cylindrical neck region, an obtriangular basal structure, and the absence of both a corona and a basal residual stalk. Its strong morphological affinities to the extant <em>Trapa natans</em> provide evidence for substantial post-Miocene evolutionary modification and divergence between European and Asian lineages, primarily influenced by regional environmental changes following the India-Asia collision. The fossil indicates that hydrologically closed basins served as refugia for <em>Trapa</em> colonization in the shallow littoral zones of freshwater lacustrine ecosystems. We infer that its proliferation was synchronous with a phase of lacustrine expansion, likely driven by enhanced Indian summer monsoon precipitation and riverine inputs into the basin. The presence of <em>T. gokarnansis</em> further indicates its survival through a climatic transition to cooler (mean temperature depression of 6–7 °C) and wetter conditions than at present. Consequently, this finding extends the documented paleobiogeographic range of <em>Trapa</em> into the eastern Himalayas and provides critical new constraints on Quaternary paleoenvironments and lacustrine ecosystem dynamics influenced by orographic uplift.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"348 ","pages":"Article 105521"},"PeriodicalIF":1.7,"publicationDate":"2026-05-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146191091","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"Systematic revision of the non-pollen palynomorph Palambages Wetzel, 1961","authors":"Facundo De Benedetti ,&nbsp;María C. Zamaloa ,&nbsp;María A. Gandolfo ,&nbsp;Néstor R. Cúneo","doi":"10.1016/j.revpalbo.2026.105501","DOIUrl":"10.1016/j.revpalbo.2026.105501","url":null,"abstract":"<div><div>The fossil non-pollen palynomorph genus <em>Palambages</em> Wetzel comprises spheroidal bodies, composed of many oval, membranous cells. The original generic diagnosis was imprecise and overly broad. Herein, its diagnosis is emended through three main modifications: 1) the number of constituent cells per cluster follows a 2ⁿ pattern; 2) clusters are exclusively hollow and comprise a single peripheral layer of thin-walled cells; and 3) each cell possesses a central operculate aperture on the outer face. The new revised description minimizes overlapping with other morphologically similar cell clusters. The type species, <em>Palambages morulosa</em>, is also emended. All six species previously attributed to <em>Palambages</em> are reassessed to evaluate their taxonomic placement. <em>P. morulosa</em> and <em>P. trilicius</em> are retained within <em>Palambages</em>. Three species are transferred to the fungal spore genera <em>Papulosporonites</em> (i.e., <em>Papulosporonites canadensis</em> comb. nov., and <em>Papulosporonites polycellularis</em> comb. nov.) and <em>Polyadosporites</em> (i.e., <em>Polyadosporites colonicus</em> comb. nov.). The species <em>P. pariunta</em> probably represents a new genus and species pending direct examination of its type material. Notably, the two species retained in the genus have been recovered from marine to brackish deposits, suggesting an affinity with marine phytoplankton—a hypothesis further supported by sedimentological and paleontological proxies.</div></div>","PeriodicalId":54488,"journal":{"name":"Review of Palaeobotany and Palynology","volume":"347 ","pages":"Article 105501"},"PeriodicalIF":1.7,"publicationDate":"2026-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145928788","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}