PalaeoworldPub Date : 2026-04-01Epub Date: 2025-12-30DOI: 10.1016/j.palwor.2025.201063
Yong-Tao Quan , Shi-Qi Wang , Hai-Yan Qi , Tong Zhang , Li-Bo Pang , Shao-Kun Chen
{"title":"First record of mammal fossil from the lower Oligocene Tala Formation, Linxia Basin, Gansu, China","authors":"Yong-Tao Quan , Shi-Qi Wang , Hai-Yan Qi , Tong Zhang , Li-Bo Pang , Shao-Kun Chen","doi":"10.1016/j.palwor.2025.201063","DOIUrl":"10.1016/j.palwor.2025.201063","url":null,"abstract":"<div><div>The Tala Formation is the bottom of the Cenozoic deposit in the Linxia Basin of Gansu Province, China, but its chronology is controversial owing to the lack of mammalian fossils. Recently, a fragmentary mandible of chalicothere <em>Schizotherium ordosium</em> was discovered from the upper part of the Tala Formation, which is the first specimen of mammalian fossil found from this formation. The fossil is assigned to <em>Schizotherium ordosium</em> based on the largest size in the genus and the metastylid separated from but smaller than the metaconid. This discovery provides a reliable reference for the paleomagnetic dating of the Tala Formation, and confirms that the age of the Tala Formation is late early Oligocene, which is meaningful for understanding the formation of the basin and sedimentary evolution.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"35 2","pages":"Article 201063"},"PeriodicalIF":1.7,"publicationDate":"2026-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146190749","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"A new tiny basal Sauropodomorpha (Dinosauria: Saurischia) from the Santa Maria Supersequence, Upper Triassic of southern Brazil","authors":"Leomir Santos Campos , Luciano Artemio Leal , Átila Augusto Stock Da-Rosa , Juliana Manso Sayão","doi":"10.1016/j.palwor.2025.201064","DOIUrl":"10.1016/j.palwor.2025.201064","url":null,"abstract":"<div><div>The Upper Triassic outcrops of Brazil and Argentina have provided important information on the earliest stages of the dinosaur radiation. Dinosaurian taxa have been increasingly discovered in these strata, contributing to the diversity of the group. Here we describe elements of dinosaurs that are among the smallest known from the Triassic of southern Brazil that show evidence of interrupted osteological growth. The material collected from the Cerro da Alemoa outcrop consists of a humerus, a metatarsal III, an ungual phalanx, a neural arch, and a vertebral centrum. Phylogenetic analysis was facilitated by anatomical comparisons, which resulted in the placement of specimen UFSM11326 within the Carnian sauropodomorph clade as a whole. The analysis of thin sections of the humerus and metatarsal III revealed a highly fragmented compact cortical structure, composed mainly of fibrolamellar bone. This was mostly composed of woven bone, surrounded by longitudinal and reticular vascularisation. The presence of an annulus and a line of arrested growth was also observed. Furthermore, the observed features suggest an individual still in development, having already undergone its first growth arrest and being midway through its second cycle. These analyses suggest ontogenetic features of a juvenile sauropodomorph with a single growth arrest. Our results provide evidence for novel growth strategies during the early evolution of this group, leading to the emergence of small-bodied dinosaurs in the Triassic.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"35 2","pages":"Article 201064"},"PeriodicalIF":1.7,"publicationDate":"2026-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146191472","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
PalaeoworldPub Date : 2026-04-01Epub Date: 2025-11-05DOI: 10.1016/j.palwor.2025.201039
Yang Yu , Qiang Li , Shi-Qi Wang , Xi-Jun Ni
{"title":"New material of Brachyhyops neimongolensis (Artiodactyla, Entelodontidae) from the upper Eocene in Jeminay, Xinjiang, China","authors":"Yang Yu , Qiang Li , Shi-Qi Wang , Xi-Jun Ni","doi":"10.1016/j.palwor.2025.201039","DOIUrl":"10.1016/j.palwor.2025.201039","url":null,"abstract":"<div><div>Entelodonts are a group of extinct artiodactyls. <em>Brachyhyops</em> is the oldest known member of the Entelodontidae, living from the late middle Eocene to late Eocene in Asia and North America. Although <em>Brachyhyops</em> is a relatively diversified genus of entelodonts in Asia, the fossil record of <em>Brachyhyops</em> is very scarce and only a few teeth and fragmentary skull and mandible pieces have been reported. Here we report a left m2 of <em>B</em>. <em>neimongolensis</em> from the upper Eocene of Xinjiang, China, which represents the first certain <em>Brachyhyops</em> fossil in Central Asia. The new material expands the distribution range of <em>B</em>. <em>neimongolensis</em> and indicates a possible tight faunal connection between Jeminay and the rest of northern China.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"35 2","pages":"Article 201039"},"PeriodicalIF":1.7,"publicationDate":"2026-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146090806","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
PalaeoworldPub Date : 2026-04-01Epub Date: 2025-11-07DOI: 10.1016/j.palwor.2025.201046
Adrisuta Ghosh , Kanishka Bose , Shiladri S. Das
{"title":"New records of Nautiloidea from the lower Eocene marine succession of the Jaisalmer Basin, western India, and their environmental implications","authors":"Adrisuta Ghosh , Kanishka Bose , Shiladri S. Das","doi":"10.1016/j.palwor.2025.201046","DOIUrl":"10.1016/j.palwor.2025.201046","url":null,"abstract":"<div><div>The lower Eocene strata of the Jaisalmer Basin have yielded a taxonomically diverse assemblage of molluscan and other invertebrate fauna. However, nautiloids remain undocumented from this region until now. The present study reports the occurrence of three nautiloid species, <em>Eutrephoceras</em> sp., <em>Angulithes rayi</em> n. sp., and <em>Angulithes</em> sp., all recovered from a single, laterally restricted stratigraphic horizon within the lower part of the Te-Takkar Member of the Khuiala Formation. The concentration of a considerable number of specimens, encompassing multiple species, is exceptional in the Cenozoic record and suggests a parautochthonous assemblage, with limited post-mortem transport.</div><div>Integrated sedimentological, petrographic, and palaeoecological evidence indicates deposition in a low-energy, shallow marine environment situated below fair-weather wave base. Biomicritic lithology, along with an associated assemblage of larger benthic foraminifera (<em>Operculina</em>, <em>Assilina</em>) and molluscs (e.g., <em>Venericardia</em>, turritelline gastropods, and chemosymbiotic bivalves), suggests warm, sub-oxic, moderately nutrient-rich conditions on soft-bottom substrates at middle-outer ramp environment. Evidence of single-flank encrustation on nautiloid shells indicates prolonged exposure at the sediment–water interface, likely under conditions of slow sedimentation and limited disturbance. Collectively, the data provide new insights into early Eocene marine ecosystems of the Jaisalmer Basin in the western Indian margin.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"35 2","pages":"Article 201046"},"PeriodicalIF":1.7,"publicationDate":"2026-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146191473","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
PalaeoworldPub Date : 2026-04-01Epub Date: 2025-11-23DOI: 10.1016/j.palwor.2025.201056
Zhi-Hao He , Xin Li , Ya-Lin Li , Atsushi Matsuoka , Hui Luo , Xiao-Hui Cui , Jian-Bo Cheng
{"title":"Early Cretaceous radiolarian assemblages in mélange of Yarlung-Tsangpo suture zone in the Saga area, southern Xizang","authors":"Zhi-Hao He , Xin Li , Ya-Lin Li , Atsushi Matsuoka , Hui Luo , Xiao-Hui Cui , Jian-Bo Cheng","doi":"10.1016/j.palwor.2025.201056","DOIUrl":"10.1016/j.palwor.2025.201056","url":null,"abstract":"<div><div>In this paper, 23 species of 14 genera within the mélange of the Saga area are documented. On this basis, a stratigraphic correlation is established between the Tethys Himalaya and the Yarlung-Tsangpo suture zone. The Cretaceous siliceous rock and siliceous mudstone strata in the mélange are also reported. Six stratigraphic units are identified in the Tethys Himalaya (Jurassic Zhela and Weimei formations, Cretaceous Rilang, Duobeng and Chuangde formations, and Paleocene Sangdanlin Formation). The Early Cretaceous radiolarians from the mélange and the deep-water sediments of the passive continental margin in the Rilang and Duobeng formations both indicate that during the Early Cretaceous, the Neo-Tethys Ocean was a widespread deep-sea environment. The reconstructed strata in this study are considered to have been deposited contemporaneously with the Rilang and Duobeng formations. In other words, while the Rilang and Duobeng formations were deposited along a passive continental margin, the deposition of the siliceous deep-water and hemipelagic sediments occurred in the ocean. After the Early Cretaceous, these sediments were preserved as accretionary mélange, becoming part of the Yarlung-Tsangpo suture zone.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"35 2","pages":"Article 201056"},"PeriodicalIF":1.7,"publicationDate":"2026-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146191323","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
PalaeoworldPub Date : 2026-04-01Epub Date: 2025-12-04DOI: 10.1016/j.palwor.2025.201058
Cícera Kelly de Morais Silva , Gabriel Ladeira Osés , Lutz Kunzmann , Domingas Maria da Conceição , William Vieira Gobo , Antonio G. Souza Filho , Cleiton C. Silva , Daniel Bernardes , Francisco Eduardo de Sousa Filho , Daniel R. do Nascimento Jr. , Maria Edenilce Peixoto Batista
{"title":"Deposition of fluorite in xylem cells of a new enigmatic seed plant from the Lower Cretaceous of Brazil","authors":"Cícera Kelly de Morais Silva , Gabriel Ladeira Osés , Lutz Kunzmann , Domingas Maria da Conceição , William Vieira Gobo , Antonio G. Souza Filho , Cleiton C. Silva , Daniel Bernardes , Francisco Eduardo de Sousa Filho , Daniel R. do Nascimento Jr. , Maria Edenilce Peixoto Batista","doi":"10.1016/j.palwor.2025.201058","DOIUrl":"10.1016/j.palwor.2025.201058","url":null,"abstract":"<div><div>The Crato Formation Flora of the Mesozoic Araripe Basin (Northeast Brazil) is characterized by a marked diversity of seed plants, among which gnetophytes and angiosperms stand out with 11 and 19 macrofossil species described, respectively. Various plant habits have been evidenced such as herbs, climbers, shrubs, and trees. Although there is a lack of taphonomic studies of plant remains from the Crato Fossil Lagerstätte, most of their reddish-brownish fossils have been assumed to have undergone a complete replacement of original organic tissues, yielding iron oxide replicas, often preserving three-dimensional cellular structures. However, the fossilization processes could be more complex than originally thought. Here, we report an intriguing and rare fossilization process in a plant fossil from this lagerstätte. The fossil is described as a new taxon <em>Nuvensia palaeobrasiliana</em> n. gen. n. sp. consisting of a sympodially organized branch with a pedunculated spike-like cone terminally attached to the medial axis and displaying helical phyllotaxis in its bracts. Tracheal elements of the axes mainly consist of tracheids with bordered pits in their radial walls and vessel elements with foraminate perforation plates. These vessel elements are scattered between the tracheids and some of them are radially organized. The limited diagnostic characters prevent affiliation within specific seed plant groups; however, the hitherto unknown combination of morpho-anatomical characters could support a relationship with gnetophytes, but without ruling out a relationship with angiosperms. Geochemical investigation of the axis revealed that the lumens of large vessel elements are filled by either calcite or fluorite, while their cell walls are preserved by iron oxides/hydroxides, which also preserve intercellular areas. The vessel elements are surrounded by silica/aluminosilicate halos. We interpret the underlying fossilization history as a sequential precipitation of calcite, fluorite, and silicates. Our results add new data to further enlighten the still not completely understood taphonomic processes leading to fluoritized plants in the fossil record. It not only provides new insights into the complex fossilization history of the usually exceptionally preserved macrofossils in the Crato Formation, but also adds a new fossil-species with remarkable xylem anatomy to the Early Cretaceous seed plants.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"35 2","pages":"Article 201058"},"PeriodicalIF":1.7,"publicationDate":"2026-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146190742","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
PalaeoworldPub Date : 2026-04-01Epub Date: 2025-11-19DOI: 10.1016/j.palwor.2025.201054
Kai Ye , Xin Rao , Quewang Danzeng , Peter W. Skelton , Bin-Tao Gao , Yi-Wei Xu , Ting Ruan , Kun Liang , Wen Guo , Ming Wang
{"title":"First record of early Aptian (Early Cretaceous) radiolitid Agriopleura libanica (Astre) from the Langshan Formation, northern Lhasa Terrane: Oldest rudist bivalves in China","authors":"Kai Ye , Xin Rao , Quewang Danzeng , Peter W. Skelton , Bin-Tao Gao , Yi-Wei Xu , Ting Ruan , Kun Liang , Wen Guo , Ming Wang","doi":"10.1016/j.palwor.2025.201054","DOIUrl":"10.1016/j.palwor.2025.201054","url":null,"abstract":"<div><div>Although the Langshan Formation has been widely regarded as spanning the Aptian to the lower Cenomanian (upper Lower to lower Upper Cretaceous), previous records of rudist bivalves from this formation were limited to the <em>Auroradiolites biconvexus</em>–<em>Magallanesia rutogensis</em> assemblage, which is restricted to upper Aptian to Albian (upper Lower Cretaceous). The discovery of <em>Agriopleura libanica</em> (Astre), along with <em>Offneria murgensis</em>, in the Langshan Formation of the northern Lhasa Terrane represents the first documented occurrence of early Aptian rudist bivalves in China, significantly extending the known stratigraphic range of rudists within this unit. The comparison between <em>A</em>. <em>libanica</em> and <em>Auroradiolites</em> supports the hypothesis that <em>Auroradiolites</em> descended directly from <em>A</em>. <em>libanica</em>, as evidenced by shared ligamentary and myophoral structures, despite differences in left valve convexity and interband morphology. The <em>A</em>. <em>libanica</em>–<em>O</em>. <em>murgensis</em> assemblage confirms the presence of a carbonate seaway in the Nyima region during the early Aptian (late Early Cretaceous), highlighting the significance of the Langshan Formation for reconstructing the paleogeography of northern Lhasa Terrane and the evolution of the Bangong-Nujiang Ocean. The occurrence of <em>A</em>. <em>libanica</em> in the Lhasa Terrane fills a critical geographic gap, bridging its known distribution from Arabia to East Asia during the early Aptian. It represents the first record of this species on the northern margin of the Tethys, as well as the first documentation of early Aptian rudists along the northern margin of the eastern Tethyan Ocean. These discoveries challenge the traditional division between northern and southern Tethyan rudist provinces, demonstrating cross-Tethyan faunal connectivity during the early Aptian.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"35 2","pages":"Article 201054"},"PeriodicalIF":1.7,"publicationDate":"2026-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146190741","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
PalaeoworldPub Date : 2026-04-01Epub Date: 2025-10-03DOI: 10.1016/j.palwor.2025.201035
Yong Yi Zhen
{"title":"Resolving a long-standing controversy in conodont taxonomy: Acodus Pander, 1856","authors":"Yong Yi Zhen","doi":"10.1016/j.palwor.2025.201035","DOIUrl":"10.1016/j.palwor.2025.201035","url":null,"abstract":"<div><div>This study documents two conodont species, <em>Acodus longibasis</em> and <em>A</em>. <em>triangularis</em>, based on a collection of over 2300 well preserved specimens from the Emanuel Formation (Lower Ordovician) of the Canning Basin, Western Australia. Both species comprise a septimembrate protoramiform — protopastinate apparatus including makellate M, protoramiform S (prototriform Sa, prototripennate Sb, protobipennate Sc and protoquadriramate Sd), and protopastinate P elements. Revision of these two species combined with a review of all known species (including the type and other form species originally named by Pander) from Europe, Australia, Asia, North America, and Argentina enables redefinition of the generic diagnosis of <em>Acodus</em>, which previously has been much debated. These new data support <em>Acodus</em> as a valid genus that phylogenetically occupies the basal position in the evolution of Prioniodontidae, a dominant and biostratigraphically important conodont family in the Ordovician.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"35 2","pages":"Article 201035"},"PeriodicalIF":1.7,"publicationDate":"2026-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146191474","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
PalaeoworldPub Date : 2026-04-01Epub Date: 2025-11-09DOI: 10.1016/j.palwor.2025.201050
Anthony J. Wright , Yong Yi Zhen , Sangmin Lee
{"title":"Early Devonian operculate rugose corals from South China and Australia","authors":"Anthony J. Wright , Yong Yi Zhen , Sangmin Lee","doi":"10.1016/j.palwor.2025.201050","DOIUrl":"10.1016/j.palwor.2025.201050","url":null,"abstract":"<div><div>The material of Early Devonian (Emsian) operculate rugose corals from Guangxi, South China, provides important morphological details leading to generic reassignment of some Chinese and Vietnamese calceolid species previously assigned to <em>Calceola</em>. Six species including two new species <em>Sinoceola gracile</em> Wright n. sp. and <em>Richtereola guangxiensis</em> Wright n. sp. and the new genus <em>Sinoceola</em> Wright n. gen. are documented. This study suggests that <em>Calceola</em> opercula are diagnostic in identification of true <em>Calceola</em>. The operculum of the diminutive <em>Sinoceola</em> n. gen. has chakeoloid major septal blades that are very different from the septa of <em>Calceola</em>, and semioval growth lines as opposed to the concentric growth lines of <em>Rhizophyllum</em> and <em>Chakeola</em>. “<em>Calceola sandalina naningensis</em>” Yoh in <span><span>Yoh and Yu, 1957</span></span> is redescribed as belonging to <em>Richtereola</em>. Opercula of <em>R</em>. <em>nanningensis</em> and <em>Richtereola guangxiensis</em> n. sp. have straight septal blades parallel to the K (counter) septum, differing from the curved septal blades of <em>Richtereola disruptus</em> <span><span>Wright, 2006</span></span>. Pragian <em>Richtereola</em> material described from New South Wales represents the oldest occurrence of the genus. Material identified as “<em>Calceola subsinensis</em> Yoh in <span><span>Yoh and Yu, 1957</span></span>” is redescribed as <em>Chakeola subsinense</em> (Yoh in <span><span>Yoh and Yu, 1957</span></span>). The holotype of <em>Calceola intermediata robusta</em> <span><span>Yoh, 1988</span></span> is considered indeterminate at genus or species level. The Vietnamese species “<em>C</em>. <em>sandalina acuminata</em>” <span><span>Mansuy, 1916</span></span> might belong to <em>Sinoceola</em> but crucial opercular details of <em>C</em>. <em>acuminata</em> are poorly known in the type material.</div><div>A new morphological term, the Extra-Alar Ridge (EAR), is proposed for the low rounded ridge on the internal surface of many calceolid corallites close to the lateral extremity of the counter face, and on the cardinal side of the alar septum where septal insertion is often seen.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"35 2","pages":"Article 201050"},"PeriodicalIF":1.7,"publicationDate":"2026-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146191324","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
PalaeoworldPub Date : 2026-04-01Epub Date: 2025-11-24DOI: 10.1016/j.palwor.2025.201057
Guang-Ying Ren , Jin-Liang Yuan , Yi-Fan Wang , Bai-Zheng An , Kai Chen , Xiao-Hui Cui , Jian Gao , Nan Li
{"title":"A new dameselloid trilobite from the Changhia Formation (Cambrian, Drumian) in central Shandong, North China","authors":"Guang-Ying Ren , Jin-Liang Yuan , Yi-Fan Wang , Bai-Zheng An , Kai Chen , Xiao-Hui Cui , Jian Gao , Nan Li","doi":"10.1016/j.palwor.2025.201057","DOIUrl":"10.1016/j.palwor.2025.201057","url":null,"abstract":"<div><div><em>Gangchengia subcylindrica</em> n. gen. n. sp., is described from the Changhia Formation (Cambrian, Drumian), central Shandong, North China. The cranidium and pygidium of the genus were predominantly found from the same stratigraphic horizon within the <em>Liopeishania lubrica</em> Zone of the upper Changhia Formation. The new genus is diagnosed by its subquadrate cranidium, broad convex subcylindrical glabella, long palpebral lobe, wide downwardly sloping preglabellar field, and narrow convex anterior border. It is also characterised by a pygidium with seven pairs of pygidial marginal spines, of which the first and sixth pairs are longer, the seventh pair is shorter.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"35 2","pages":"Article 201057"},"PeriodicalIF":1.7,"publicationDate":"2026-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146191470","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}