{"title":"Dental morphology of the pycnodontid fish †Stemmatodus rhombus (Agassiz 1844) (Neopterygii, †Pycnodontiformes) from the Early Cretaceous, with comments on its systematic position","authors":"J. Kriwet","doi":"10.1017/S0263593300000560","DOIUrl":"https://doi.org/10.1017/S0263593300000560","url":null,"abstract":"ABSTRACT The dental morphology of the pycnodontid fish †Stemmatodus rhombus (Agassiz, 1844) from the Lower Cretaceous (lower Barremian) of southern Italy is described in detail. SEM pictures of vomerine and prearticular teeth are provided for the first time allowing the comparison with other Early Cretaceous pycnodonts, which are only known from isolated dental remains. The dentition of †Stemmatodus rhombus consists of incisiform grasping teeth on the paired premaxillae and dentaries and crushing teeth on the unpaired vomer and paired prearticulars. The vomerine teeth are arranged in five longitudinal rows, which display symmetry with a distinct median principal and two lateral rows. The prearticular bears three longitudinal rows. Teeth of the main row are much wider than those of the lateral ones. The tooth crowns are rather flat and expose a shallow apical depression, which is surrounded by a ring of widely-spaced small tubercles. The ornamentation and arrangement of teeth distinguishes dentitions of †Stemmatodus from all other pycnodonts. The wear pattern of the teeth is evaluated and the significance of the dental morphology in pycnodont taxonomy is discussed. In addition, the systematic position of †Stemmatodus is discussed.","PeriodicalId":83368,"journal":{"name":"Transactions of the Royal Society of Edinburgh: earth sciences","volume":"30 1","pages":"145 - 155"},"PeriodicalIF":0.0,"publicationDate":"2003-06-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"84550401","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Lithofacies and sedimentary cycles within the Late Dinantian (late Brigantian) of Fife and East Lothian: is a sequence stratigraphical approach valid?","authors":"A. M. Kassi, J. Weir, J. Mcmanus, M. Browne","doi":"10.1017/S0263593300000547","DOIUrl":"https://doi.org/10.1017/S0263593300000547","url":null,"abstract":"ABSTRACT The late Brigantian topmost parts of the Pathhead Formation (Aberlady Formation in East Lothian) and the succeeding Lower Limestone Formation crop out widely in Fife and East Lothian. The successions include nine deltaic, coastal floodplain and marine shelf cycles (cyclothems), of which the lowest examined terminates the Pathhead and Aberlady Formations and the remaining eight constitute the Lower Limestone Formation. The cyclothems conform broadly to the ‘Yoredale” transgressive/regressive pattern in which a transgressive marine shelf phase is succeeded by delta progradation and terminates with a fluvial delta plain phase. Cycles may combine to form compound cyclothems up to more than 50 m thick, in which a basal, typically complete initial cycle of Yoredale pattern is succeeded by up to five base-absent minor cycles. These are thinner, more variable and less laterally persistent units in which the marine phase is weakly represented or absent. Cyclothems reflect successive marine flooding events, possibly under eustatic control, succeeded by delta progradation and, ultimately, leading to extensive palaeosol formation, including coal seams. Sedimentation and palaeosol formation were partly controlled by fault-induced differential subsidence and are likely to have been related to autocyclic processes. Local uplift and subsidence associated with vulcanicity, as at Kinghorn and Elie, have led to thickening or thinning of sediments accumulated in a given time period. Initial cycles initiate longer-period allocycles, corresponding broadly to third-order Exxon Production & Research (EPR) Type 1 sequences having a periodicity of around 1 Ma, within the Milankovitch orbital band. Two parasequences constitute each initial cycle: a lower, initiated on a marine flooding surface, and an upper, bounded by the base of the lowest thick sandstone in the cycle; cyclothem bases and sequence bases thus alternate. Parasequences and sequences are less well defined in minor cycles due to the problem of tracing the combined disconformity and soil profile of the underclay beyond the edge of channel sandstones. Minor cycles were controlled primarily by short-period autocyclic sedimentary and, or, tectonic processes, including delta-lobe switching and differential subsidence. Although we have attempted to interpret the deposits of Fife and the Lothians in terms of sequence stratigraphy, we are not fully convinced that the patterns of associated changes widely recognised within the framework of sequence stratigraphy can be confidently applied in succesions in which autocyclic changes feature strongly in an area undergoing active basin subsidence associated with strike-slip faulting. There is no doubt that some of the cyclicity discerned in the late Brigantian successions of eastern Scotland was related to eustatic sea level changes, which gave rise to the widespread limestone platforms or marine bands. The formation of eight cyclothems within the 2·5–3·5 Ma of lat","PeriodicalId":83368,"journal":{"name":"Transactions of the Royal Society of Edinburgh: earth sciences","volume":"23 1","pages":"95 - 113"},"PeriodicalIF":0.0,"publicationDate":"2003-06-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"81006078","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"A marginal marine ichnofauna from the Blaiklock Glacier Group (?Lower Ordovician) of the Shackleton Range, Antarctica","authors":"B. Weber, S. Braddy","doi":"10.1017/S026359330000050X","DOIUrl":"https://doi.org/10.1017/S026359330000050X","url":null,"abstract":"ABSTRACT A diverse arthropod-dominated ichnofauna, associated with a poorly preserved crustacean fauna and soft-bodied ?medusoid impressions, is described from the Blaiklock Glacier Group of the north-western Shackleton Range (Coats Land), Antarctica. The ichnofauna consists of Asaphoidichnus, Beaconites, Didymaulichnus, Diplichnites, Gordia, ?Laevicyclus, Merostomichnites, Monomorphichnus, Palaeophycus, Planolites, Rusophycus, Selenichnites, and Taphrhelminthoides (ichnogen nov.). Three new ichnotaxa are recognised: Taphrhelminthoides antarcticus n. ichnogen. et ichnosp. is a bilobate trail, composed of two parallel flat lobes, separated by a median ridge with a characteristic figure-of-eight pattern. Merostomichnites gracilis n. ichnosp. is characterised by its proportions (external:internal width ratio >3) and series of 10 to 12, thin, linear tracks. Selenichnites antarcticus n. ichnosp. is characterised by small elongate horseshoe-shaped marks, the medial portion showing three to five transverse scratch-marks. The palaeoenvironment is interpreted as extremely shallow marine water, possibly a tide-dominated estuary, based on sedimentological evidence and the composition of the ichnofauna. Radiometric and palaeomagnetic data indicate that this assemblage is Lower Ordovician in age, representing the first autochthonous Ordovician fossiliferous succession to be described from Antarctica. The succession shows several sedimentological and palaeontological similarities with the basal units of the Ordovician Table Mountain Group in South Africa, supporting palaeogeographic models placing the Palaeozoic Blaiklock basin close to the Ordovician Table Mountain basin.","PeriodicalId":83368,"journal":{"name":"Transactions of the Royal Society of Edinburgh: earth sciences","volume":"19 1","pages":"1 - 20"},"PeriodicalIF":0.0,"publicationDate":"2003-03-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"89442669","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"The skull of Cochleosaurus bohemicus Frič, a temnospondyl from the Czech Republic (Upper Carboniferous) and cochleosaurid interrelationships","authors":"S. Sequeira","doi":"10.1017/S0263593300000511","DOIUrl":"https://doi.org/10.1017/S0263593300000511","url":null,"abstract":"ABSTRACT The cranial anatomy of the primitive Carboniferous temnospondyl Cochleosaurus bohemicus is described herein from several large, presumably adult, skulls. This new morphological data modifies earlier descriptions by Frič (=Fritsch in post-1876 publications), Romer and Steen, which were based almost entirely on small, subadult Cochleosaurus bohemicus specimens. Recognition of a new autapomorphy for Cochleosaurus bohemicus, (pitting on the ventral surface of the postorbitals) which is independent of ontogenetic change, has facilitated more accurate intra-specific comparisons between cochleosaurids. Cladistic analysis reveals a clear dichotomy separating the Edopoidea from the more derived clade of the remaining temnospondyls, which includes Balanerpeton and Dendrerpeton. The Edopoidea are only weakly supported as a superfamily in this analysis. The Cochleosauridae are a monophyletic family comprising the genera Procochleosaurus, Adamanterpeton, Cochleosaurus and Chenoprosopus.","PeriodicalId":83368,"journal":{"name":"Transactions of the Royal Society of Edinburgh: earth sciences","volume":"28 1","pages":"21 - 43"},"PeriodicalIF":0.0,"publicationDate":"2003-03-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"78854189","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"The seymouriamorph tetrapod Utegenia shpinari from the ?Upper Carboniferous–Lower Permian of Kazakhstan. Part I: Cranial anatomy and ontogeny","authors":"Jozef Klembara, M. Ruta","doi":"10.1017/S0263593300000523","DOIUrl":"https://doi.org/10.1017/S0263593300000523","url":null,"abstract":"ABSTRACT The skull roof, palate and lower jaw of the seymouriamorph Utegenia shpinari (?Upper Carboniferous–Lower Permian; Kurgalin Formation, Kazakhstan) are described in detail, and a new cranial reconstruction is presented. Important features include: skull slightly longer than broad; elongate nasals; deep, elongate posterior cheek region; long, dorsoventrally deep posterior jugal ramus; flat, recurved, blade-like tabular process; denticle-covered anterior ventral part of pterygoid quadrate ramus; transverse pterygoid flange; low radiating ridges with denticle rows on pterygoid and parasphenoid cultriform process; incipient anterior wedge-like process on posterior end of parasphenoid cultriform process; distinct, robust posterolateral processes of posterior plate of parasphenoid. Degree of skull ossification, ornamentation, and absence of ossified quadrate, articular and endocranium suggest that large Utegenia specimens correspond to larval Discosauriscus. In such specimens, dentine infolding extends for almost half of the crown in premaxillary and largest maxillary teeth, as in late larval/metamorphic Discosauriscus. Preliminary results of a regression analysis through some simple skull measurements indicates that, although the preorbital and interorbital regions elongate isometrically relative to the posterior skull table, they do not show a particularly strong linear correlation with the widening of the skull during ontogeny.","PeriodicalId":83368,"journal":{"name":"Transactions of the Royal Society of Edinburgh: earth sciences","volume":"44 1","pages":"45 - 74"},"PeriodicalIF":0.0,"publicationDate":"2003-03-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"90541800","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"The seymouriamorph tetrapod Utegenia shpinari from the ?Upper Carboniferous–Lower Permian of Kazakhstan. Part II: Postcranial anatomy and relationships","authors":"Jozef Klembara, M. Ruta","doi":"10.1017/S0263593300000535","DOIUrl":"https://doi.org/10.1017/S0263593300000535","url":null,"abstract":"ABSTRACT The postcranial skeleton of the seymouriamorph Utegenia shpinari (?Upper Carboniferous–Lower Permian, Kurgalin Formation, Kazakhstan) is redescribed. Features that distinguish it from other Lower Permian seymouriamorphs (Discosauriscus, Ariekanerpeton, Seymouria) are: broad anterior portion of the interclavicle stem, merging indistinctly into the interclavicle plate; absence of a bulge in the anterior half of such a stem; presence of at least 28 presacral vertebrae; gastralia. The poorly ossified limbs of the largest Utegenia specimens are similar in degree of development to those of larval Discosauriscus, but the almost cylindrical, anterior trunk pleurocentra recall the condition of early juvenile Discosauriscus. A phylogeny of the best known seymouriamorphs, using a small but diverse exemplar from other early tetrapod groups, places Utegenia within seymouriamorphs as sister taxon to discosauriscids (Ariekanerpeton plus Discosauriscus). This conclusion affects origin and dispersal scenarios for seymouriamorphs, and supports the hypothesis of a widespread geographical record of Lower Permian taxa spanning across western Euramerica and eastern Asia.","PeriodicalId":83368,"journal":{"name":"Transactions of the Royal Society of Edinburgh: earth sciences","volume":"86 1","pages":"75 - 93"},"PeriodicalIF":0.0,"publicationDate":"2003-03-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"89040491","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"A revised correlation of Silurian rocks in the Girvan district, SW Scotland","authors":"J. Floyd, M. Williams","doi":"10.1017/S0263593300000481","DOIUrl":"https://doi.org/10.1017/S0263593300000481","url":null,"abstract":"ABSTRACT In late Ordovician and early Silurian times, the Girvan district lay in a shelf marinesetting on the margin of Laurentia, on the northern side of the Iapetus Ocean. The Lower Palaeozoic rocks of the Girvan district, and their shelly and graptolitic fossil fauna, were systematically described by Lapworth in 1882 and have formed an important research resource ever since. They provide valuable evidence for the depositional environment and geological setting of Girvan during the early Palaeozoic, in both regional and wider contexts, and demonstrate the long-recognised close affinity with contemporaneous Laurentian faunas. However, by late Ordovician and into Silurian times, the earlier Iapetus oceanic barrier to faunal migration had largely gone and there is good correlation between contemporaneous marine fauna throughout the British Isles and Scandinavia. Despite much recent research in the area, including resurvey work by the British Geological Survey, no comprehensive review of Silurian lithostratigraphy at Girvan has been published since the revision by Cocks and Toghill in 1973. The present review of the Silurian rocks addresses this need and complements the recently published (Fortey et al. 2000) revision of the underlying Ordovician rocks, thus bringing the entire Girvan Lower Palaeozoic succession up to modern standards of nomenclature.","PeriodicalId":83368,"journal":{"name":"Transactions of the Royal Society of Edinburgh: earth sciences","volume":"33 1","pages":"383 - 392"},"PeriodicalIF":0.0,"publicationDate":"2002-12-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"89901201","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Adequacy of the Early Ordovician trilobite record in the southern Montagne Noire (France): biases for biodiversity documentation","authors":"D. Vizcaïno, J. Álvaro","doi":"10.1017/S0263593300000493","DOIUrl":"https://doi.org/10.1017/S0263593300000493","url":null,"abstract":"ABSTRACT The litho- and biostratigraphical subdivisions of the Tremadocian-Arenigian succession of the southern Montagne Noire are hereby revised. The Lower Ordovician diversity patterns are estimated through statistical analysis of 27 families, 64 genera and 132 species of trilobites across nine interval zones; from bottom to top: the Proteuloma geinitzi, Shumardia (Conophrys) pusilla, Euloma filacovi, Taihungshania miqueli, Taihungshania shui landyranensis, Colpocoryphe maynardensis, Neseuretus (Neseuretus) arenosus, Apatokephalus incisus and Hangchungolithus primitivus zones. Maximum trilobite diversity occurred within the E. filacovi and A. incisus zones, alternating with two sharp declines (S. (C.) pusilla and T.shui landyranensis zones), in both cases drastically related to transgressive-regressive trends. Trilobites attained their highest diversity in muddy outer-platform settings, decreasing in fine-grained sandstones and siltstones of shoreface environments, where trilobites could still be numerically abundant. Four major trilobite turnovers are recognised at the bottom of the P. geinitzi, S.(C.)pusilla, E. filacovi and T. shui landyranensis zones, in which survivors from previous intervals are not reported at specific level, and important generic replacements took place. Despite the relative incompleteness of the Lower Ordovician trilobite record, envisaged after analysing the high proportion of monotypic taxa and discontinuous ranges of some families, the trilobite record is representative enough to estimate diversity patterns, although a better understanding of the palaeoenvironmental control will greatly enhance biodiversity resolution.","PeriodicalId":83368,"journal":{"name":"Transactions of the Royal Society of Edinburgh: earth sciences","volume":"10 6 1","pages":"393 - 401"},"PeriodicalIF":0.0,"publicationDate":"2002-12-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"83247455","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Evolutionary relationships of the Sparidae (Teleostei: Percoidei): integrating fossil and Recent data","authors":"J. J. Day","doi":"10.1017/S0263593300000468","DOIUrl":"https://doi.org/10.1017/S0263593300000468","url":null,"abstract":"ABSTRACT The Eocene sparid fauna (Teleostei: Percoidei) from Monte Bolca, Italy and from the London Clay, U.K. is revised based on re-examination of the type material and phylogenetic analyses of primarily osteological data. Two phylogenetic analyses, one of the Eocene taxa and a combined analysis of fossil and extant taxa, were performed. The addition of fossils to the extant data greatly increased numbers of most parsimonious trees, destabilising and obscuring basal relationships within the Sparidae. Combination of the data from fossil and extant data also affected relationships among the fossil taxa, changing some from those recovered using fossil data alone and destabilising others. Successive approximations character weighting supported the inclusion of the Eocene taxa within a monophyletic Sparidae. The genus Sparnodus, as previously conceived, is paraphyletic and is partitioned to remove the paraphyly. Five monotypic genera are recognised, including three new genera, Abromasta, Ellaserrata and Pseudosparnodus. Inclusion of the fossils in the phylogenetic analysis implies a minimum age of origin for the Sparidae of 55 Ma with most Recent sparid fauna in place no later than the Miocene, and provides further evidence that the diversification of feeding strategies occurred early on in the evolutionary history of the group.","PeriodicalId":83368,"journal":{"name":"Transactions of the Royal Society of Edinburgh: earth sciences","volume":"134 1","pages":"333 - 353"},"PeriodicalIF":0.0,"publicationDate":"2002-12-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"77387697","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"A new crustacean from the Early Devonian Rhynie chert, Aberdeenshire, Scotland","authors":"S. Fayers, N. Trewin","doi":"10.1017/S026359330000047X","DOIUrl":"https://doi.org/10.1017/S026359330000047X","url":null,"abstract":"ABSTRACT A new crustacean, Castracollis wilsonae is described from a loose block of the Early Devonian Rhynie chert, found in the vicinity of Rhynie, Aberdeenshire, Scotland. It differs markedly from Lepidocaris rhyniensis, Scourfield 1926, the only other crustacean found in the Rhynie chert. The material comprises complete individuals up to 8 mm long and fragmentary remains. The head is normally damaged and detached from the body, and damage to the anterior of the trunk suggests the specimens are exuviae. The head appears domed with a labrum, robust mandibles, and long biramous antennae. A few specimens exhibit a probable detached cephalo-thoracicshield. The trunk is multi-segmented, comprising similar ring-like somites. The thorax variably comprises up to 26 segments. The anterior 11 segments possess similar long, phyllopodous appendages, the remainder variably possess 10 to 15 phyllopodous appendages with a disposition of one per somite. The abdomen comprises up to 28 apodous segments. The posterior of the body comprises a telson with two furcal rami. The crustacean is most probably a calmanostracan branchiopod. It occurs associated with Lepidocaris, charophytes, cyanobacteria and coprolites within a ‘clotted’ chert texture, indicating subaqueous deposition, most probably in a temporary freshwater pool in an area of surficial hydrothermal activity.","PeriodicalId":83368,"journal":{"name":"Transactions of the Royal Society of Edinburgh: earth sciences","volume":"68 1","pages":"355 - 382"},"PeriodicalIF":0.0,"publicationDate":"2002-12-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"76296847","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}