Journal of Systematic Palaeontology最新文献

{"title":"Phylogenetic analysis of the gastropod genus Calliotropis Seguenza, 1902 (Vetigastropoda: Calliotropidae), including fossil and living species","authors":"D. Pérez, Mariel Ferrari, M. Ezcurra","doi":"10.1080/14772019.2022.2100288","DOIUrl":"https://doi.org/10.1080/14772019.2022.2100288","url":null,"abstract":"The taxonomic content of the genus Calliotropis and the systematic positions of the fossil species referred to this genus have been under debate because of the striking resemblance they bear to their extant counterparts in terms of general shell morphology and ornament pattern. In a recent contribution, two subgenera of Calliotropis were preliminarily distinguished and considered to be a result of ecomorphological variation translated into evolutionary changes: shallow-reef ecomorphotypes, representing the fossil species of Calliotropis (Riselloidea), and deep-water ecomorphotypes, grouping the living forms of Calliotropis (Calliotropis). However, this division of Calliotropis species was based on qualitative grounds, lacking a quantitative phylogenetic analysis sampling both modern and fossil taxa. The present research aims to comprehensively revisit this systematic scheme after a morphological delimitation of calliotropid lineages. We use a quantitative phylogenetic analysis to shed light on the evolutionary history of this group of gastropods. Our results indicate that the Mesozoic genus ‘Calliotropis’ is not monophyletic, but living species comprise a monophyletic group with a Pliocene–Recent distribution. Fossil ‘Calliotropis’ species represent a paraphyletic assemblage composed of different clades: the monophyletic Riselloidea (Cossmann) including only Jurassic species; a Palaeogene group represented by the new genus Arachnotropis gen. nov.; and a monophyletic Triassic group that matches the content of the previously described genus Trochonodus Nützel et al., 2003. The genus Ambercyclus Ferrari et al., 2014 is monophyletic and also related to the above-mentioned taxa. http://zoobank.org/urn:lsid:zoobank.org:pub:0BF29053-3D67-41A4-923D-5621D448AD61","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":"1 - 18"},"PeriodicalIF":2.6,"publicationDate":"2022-08-24","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"43532556","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"Unravelling the identity of the platanistoid Notocetus vanbenedeni Moreno, 1892 (Cetacea, Odontoceti) from the early Miocene of Patagonia (Argentina)","authors":"M. Viglino, M. Buono, Yoshihiro Tanaka, J. Cuitiño, R. Fordyce","doi":"10.1080/14772019.2022.2082890","DOIUrl":"https://doi.org/10.1080/14772019.2022.2082890","url":null,"abstract":"Platanistoidea remains one of the most evolutionarily intriguing lineages of toothed whales (Odontoceti). The clade comprises mostly extinct species from the late Oligocene–early Miocene onward and a single extant riverine genus (Platanista). There is an ongoing debate as to the membership of Platanistoidea and the causes of their near extinction. In Patagonia (Argentina), the most abundant platanistoid recorded in the lower Miocene Gaiman Formation is Notocetus vanbenedeni, first described by Moreno in 1892 based on two individuals. The goal of the present contribution is to conduct an updated anatomical, palaeobiological and phylogenetic analyses of Notocetus vanbenedeni and hence contribute to an understanding of the evolutionary history of the Platanistoidea. Our analyses, including at least 26 individuals (12 undescribed), show that Notocetus vanbenedeni is a valid platanistoid taxon, recovered as part of a new clade. Among its most outstanding features, this taxon has an elevated dorsal tubercular supraorbital crest formed mainly by the frontal, the precursor of the pneumatized crest of the extant Platanista. Notocetus vanbenedeni also shows initial stages of the plesiomorphic bony connection between the earbones and skull as in Platanista, although the functional implications for hearing remain elusive. The nasal sac system, pterygoid sinus system and morphology of the earbones suggest that this species was able to hear high-frequency sounds and echolocate underwater, similar to extant odontocetes. Thus, Notocetus vanbenedeni presents a mosaic of features that suggest an intermediate platanistoid morphotype. Anatomical differences and phylogenetic analyses suggest that Peruvian specimens could not be referred to this species. The feeding apparatus of Notocetus vanbenedeni makes it the only combination suction-feeder recorded in the early Miocene of Patagonia and among the smallest odontocetes. Finally, the abundant records of Notocetus vanbenedeni in an inner shelf environment with freshwater influence suggest a possible early preference for such protected habitats.","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":" ","pages":""},"PeriodicalIF":2.6,"publicationDate":"2022-08-24","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"44131506","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"Towards a unifying systematic scheme of fossil and living billfishes (Teleostei, Istiophoridae)","authors":"Carlos De Gracia, A. Correa-Metrio, M. Carvalho, J. Vélez‐Juarbe, T. Přikryl, C. Jaramillo, J. Kriwet","doi":"10.1080/14772019.2022.2091959","DOIUrl":"https://doi.org/10.1080/14772019.2022.2091959","url":null,"abstract":"Extant istiophorids are open ocean apex predators that are extensively studied due to their ecological importance and high values for fisheries. Nevertheless, little is known about their evolution because of a fragmentary fossil record and extremely difficult taxonomy of fossil species. Here, we present a new phylogenetic hypothesis covering fossil and living istiophorids. Our results demonstrate that istiophorid richness is larger than previously assumed, comprising eight genera with 20 species. The phylogenetic analysis shows that istiophorids are grouped into four clades: the Istiophorus clade, which includes the sailfish; the Machairostra clade, which comprises Makaira spp., including two new species from the late Miocene (†Makaira colonense sp. nov. and †Makaira fierstini sp. nov.); the Gracilorostra clade, which comprise all remaining istiophorids with exception of spearfishes and includes two new genera and one new species (†Morgula donosochagrense gen. et sp. nov. and †Spathochoira calvertense gen. et. comb. nov.); and the Tetrapturomorpha clade is composed of the spearfishes and the extinct †Prototetrapturus courcelli gen. et. comb. nov. The family Istiophoridae shows an evolutionary trend toward reduction of the premaxillary thickness and increasing the extension of narial cavities. This reduction is related to an increase of adipose tissues in the rostrum base probably driven by the presence of the oleofera gland, an organ involved in feeding, healing, endothermy and hydrophobic functions. Our phylogeny shows a direct relationship between the rostral and cranial shape explained by body size and feeding behaviour. The larger istiophorids have lateral apophysis and the larger spines of the vertebral column. The spearfishes represent the smaller species of the family, with the extant Tetrapturus spp. first appearing in the late Pliocene. The clade Tetrapturomorpha shows an extreme size reduction over time when compared with species of their sister clade Gracilorostra, demonstrating an evolutionary trend towards size reduction. http://zoobank.org/urn:lsid:zoobank.org:pub:D3D3B15B-36FA-42EB-98AD-FAF369D989EB","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":"1 - 36"},"PeriodicalIF":2.6,"publicationDate":"2022-08-19","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"46360902","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"New allotherian specimens from the Middle Jurassic Woodeaton Quarry (Oxfordshire) and implications for haramiyidan diversity and phylogeny","authors":"Fangyuan Mao, Philippa Brewer, J. Hooker, Jin Meng","doi":"10.1080/14772019.2022.2097021","DOIUrl":"https://doi.org/10.1080/14772019.2022.2097021","url":null,"abstract":"We report new allotherian tooth specimens from the Middle Jurassic White Limestone Formation at Woodeaton Quarry (Oxfordshire), United Kingdom. Two teeth are assigned to Kermackodon (=Eleutherodon) oxfordensis, a taxon whose original generic name (Eleutherodon) was preoccupied and is here assigned to Kermackodon to form a new binomial combination for the species name. Butlerodon quadratus gen. et sp. nov. (family Kermackodontidae), based on 13 cheek teeth and incisors, shows dental features intermediate between K. oxfordensis and the Late Triassic “haramiyidans” (Haramiyavia and Thomasia). Woodeatonia parva gen. et sp. nov. (family indeterminate), based on three teeth, is characterized by its small size. A second upper molar from a multituberculate is identified as Hahnotherium cf. H. antiquum, which possesses characters typical for multituberculates but distinctive from “haramiyidans”. The allotherian teeth from the Forest Marble Formation, previously assigned to the haramiyidans “Eleutherodon”, “Millsodon” and “Kirtlingtonia”, and the multituberculate Kermackodon, are reinterpreted as teeth from different upper or lower dental loci of the same haramiyidan species K. oxfordensis, which result in significant taxonomical modification of these allotherians (“haramiyidans” and multituberculates). Given that Kermackodon has been regarded as a transitional form between multituberculates and “haramiyidans”, these taxonomical modifications would affect interpretation of early evolution of allotherians. In a comparison of molars in known “haramiyidans”, we delve into their occlusal patterns and cusp homologies that have been controversial but pivotal for understanding evolution of allotherians. We further conduct the first phylogenetic analysis of haramiyidan species. The European Late Triassic species form the stem-ward taxa of “haramiyidans” and the Jurassic species from the United Kingdom are grouped with arboroharamiyids from the Yanliao Biota, China, and nested in “haramiyidans”. http://zoobank.org/urn:lsid:zoobank.org:pub:38F1FB5A-17A2-498F-B1FD-38D975548201 http://zoobank.org/urn:lsid:zoobank.org:act:FF5ADA10-C044-4541-ADA6-54448F17C673 http://zoobank.org/urn:lsid:zoobank.org:act:6E59071A-56DF-43D2-A140-2860465BFECA http://zoobank.org/urn:lsid:zoobank.org:act:CA2EE3CF-951C-450A-879D-2AE438F6BB2A http://zoobank.org/urn:lsid:zoobank.org:act:6B26D9C3-9FC3-4517-A1EF-51822067F970","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":"1 - 37"},"PeriodicalIF":2.6,"publicationDate":"2022-08-19","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"42019091","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"New rodents shed light on the age and ecology of late Miocene ape locality of Tapar (Gujarat, India)","authors":"R. Patnaik, N. P. Singh, K. M. Sharma, N. A. Singh, Deepak Choudhary, Y. P. Singh, Rohit Kumar, W. A. Wazir, A. Sahni","doi":"10.1080/14772019.2022.2084701","DOIUrl":"https://doi.org/10.1080/14772019.2022.2084701","url":null,"abstract":"The Miocene ape (Sivapithecus) locality of Tapar in Kutch (Gujarat, India) has yielded a diverse rodent assemblage that includes: a new murine Progonomys prasadi sp. nov., a new gerbilline Myocricetodon gujaratensis sp. nov., a new rhizomyne Kanisamys kutchensis sp. nov. and a new sciurine Tamias gilaharee sp. nov., beside additional remains of Progonomys morganae, Dakkamys asiaticus, Prokanisamys sp., Sayimys sivalensis and Democricetodon fejfari. Morphometric and PAUP based phylogenetic analyses place Progonomys prasadi sp. nov. within the Progonomys lineage. The cladogram obtained for the Siwalik murines suggest that Progonomys was ancestral to all the modern and one extinct murine genera recovered from the Siwaliks. The advanced features of Myocricetodon gujaratensis sp. nov. indicate that it was an immigrant to the subcontinent in the late Miocene. The cladistic analysis performed on Kanisamys kutchensis sp. nov. shows that it shared several advanced characters with contemporaneous Kanisamys nagrii and Kanisamys sivalensis. Based on the biostratigraphical ranges of Siwalik rodents and the co-occurrence of advanced forms of new and already reported murines, a new gerbilline and a new sciurine, we propose an age of ∼10 Ma to the primate-bearing Tapar locality. Already reported stable isotope data on murines, and ecological preferences of modern counterparts of the fossil rodents and associated sharks and rays from Tapar locality, indicate that the Miocene ape Sivapithecus may have lived in a subtropical monsoonal forest close to the coast, very different from the present day arid conditions. http://zoobank.org/urn:lsid:zoobank.org:pub:12CE1B44-22A0-450F-9588-6C7F25242771","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":" ","pages":""},"PeriodicalIF":2.6,"publicationDate":"2022-08-05","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"48387038","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"A new podocnemidid (Pleurodira: Pelomedusoides) from the Eocene of north-western Argentina, with comments on its evolutionary relationships and palaeoenvironmental settings","authors":"M. S. de la Fuente, Ignacio J. Maniel, Pedro Parraguez Ruiz, J. Ledesma, M. V. Deraco, C. D. del Papa, C. Herrera","doi":"10.1080/14772019.2022.2081939","DOIUrl":"https://doi.org/10.1080/14772019.2022.2081939","url":null,"abstract":"We report a new small podocnemidid, Gestemys powelli gen. et sp. nov., based on one nearly complete skull, lower jaw, two shells, and remains of the appendicular skeleton belonging to two specimens from the Eocene Geste Formation of the San Antonio de la Cobres Basin, Salta Province, north-western Argentina. As in other podocnemidid genera, Gestemys exhibits a fully developed and medially extensive cavum pterygoidei, an incisura columellae auris enclosing the stapes and Eustachian tube, and lacks an exoccipital quadrate contact. Gestemys powelli bears a huge foramen palatinum posterius and a short pterygoid flange that does not reach the basisphenoid suture, exposing the cavum pterygoid. Phylogenetic analyses place Gestemys powelli as a member of Podocnemididae more closely related to Erymnochelyinae than Podocnemidinae. In regards to the palaeoenvironmental settings of the Palaeogene formations (Maíz Gordo and Geste) of north-western Argentina, at least two features differentiate the palaeoenvironment where G. powelli was recovered and those of other podocnemidid turtles from the Maíz Gordo Formation: 1) G. powelli was found in situ with very little re-working; and 2) in contrast to the muddy rocks indicating the marginal lacustrine setting and humid climate of the Maíz Gordo Formation, G. powelli is associated with a coarse-grained floodplain setting and a dry, temperate climate. Such differences highlight the contrasting environmental dynamics in which podocnemidid turtles lived during the Palaeogene. http://zoobank.org/urn:lsid:zoobank.org:pub:0E631DA2-053F-4BD6-91BD-2401F34A5777","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":""},"PeriodicalIF":2.6,"publicationDate":"2022-07-25","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"43519976","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"A comparative and correlational study of the Vocal Fatigue Index (VFI) and the Voice Fatigue Handicap Questionnaire (VFHQ) in individuals with and without voice disorders.","authors":"Lingyu Yu, Jia Ren, Ting Xu, Tianpei Ma, Zhongjing Pan, Dan Lu","doi":"10.1016/j.jvoice.2022.04.012","DOIUrl":"10.1016/j.jvoice.2022.04.012","url":null,"abstract":"<p><strong>Objectives: </strong>The aim of this study was to examine the difference and correlation between the Vocal Fatigue Index (VFI) and the Voice Fatigue Handicap Questionnaire (VFHQ) among individuals with and without voice disorders.</p><p><strong>Study design: </strong>Observational study.</p><p><strong>Methods: </strong>This study was performed from June to December 2021. A total of 308 people were enrolled, including 72 men and 236 women. Questionnaire scores were obtained for both the VFI and the VFHQ. The Mann-Whitney U test and Spearman correlational analysis were used to analyze the data.</p><p><strong>Results: </strong>Of the 308 participants, 186 had a voice disorder and 122 did not. The most common voice disorders were chronic laryngitis and vocal fold polyps. The VFI and VFHQ scores differed between groups with and without voice disorders and between men and women. The VFHQ total score correlated significantly with all subscale scores of the VFHQ and with the VFI factor scores.</p><p><strong>Conclusions: </strong>People with voice disorders have significantly worse voice-related quality of life and more severe vocal fatigue symptoms than those without. The scores on the VFI and VFHQ correlate significantly.</p>","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"4 1","pages":""},"PeriodicalIF":2.2,"publicationDate":"2022-05-21","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"79038279","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"Phylogenetic analysis of Ceratophryidae (Anura: Hyloidea) including extant and extinct species","authors":"L. A. Barcelos, Diego Almeida-Silva, C. M. D. Santos, V. K. Verdade","doi":"10.1080/14772019.2022.2050824","DOIUrl":"https://doi.org/10.1080/14772019.2022.2050824","url":null,"abstract":"The Neotropical frog family Ceratophryidae is composed of wide-mouthed frogs with stout bodies. Living species of the family are consistently recovered as a monophyletic group, but with disparities among analyses regarding internal relationships. Ceratophryidae presents one of the richest fossil records in Anura. Nevertheless, phylogenetic analyses including both extant and extinct species are still scarce, and the position of fossils is persistently debated. In this sense, the systematics of the family has changed considerably in the last decade with the exclusion of Baurubatrachus pricei (Late Cretaceous), Beelzebufo ampinga (Late Cretaceous) and Wawelia gerholdi (early Miocene). Herein, a morphologically based phylogeny for Ceratophryidae, including living species (11 spp.) and fossil specimens (10 spp.), is used as a background to discuss the evolutionary history of the family and its classification. We phylogenetically placed Baurubatrachus pricei, Beelzebufo ampinga and Wawelia gerholdi as non-ceratophryids. We recovered a monophyletic Ceratophryidae: Lepidobatrachus and Ceratophrys form a clade, with Chacophrys as its sister group. Our analysis corroborates the C. cornuta and C. aurita groups. Among fossils, L. australis and C. sagani were recovered as valid species based on autapomorphies, and C. rusconii was found to be the sister of all Ceratophrys. Ceratophrys ensenadensis, C. ameghinorum, C. aurita NHMUK PV OR18895/6 and C. sagani belong to the C. aurita group. We also discuss homoplasies in Ceratophryidae, divergence-time estimates, and the evolution of ploidy and a dorsal shield in the family.","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"19 1","pages":"1449 - 1466"},"PeriodicalIF":2.6,"publicationDate":"2022-05-04","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"43411251","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"The Ordovician diversification of sea urchins: systematics of the Bothriocidaroida (Echinodermata: Echinoidea)","authors":"J. Thompson, L. Cotton, Y. Candela, M. Kutscher, M. Reich, D. Bottjer","doi":"10.1080/14772019.2022.2042408","DOIUrl":"https://doi.org/10.1080/14772019.2022.2042408","url":null,"abstract":"The echinoids of the order Bothriocidaroida represent the initial burst of sea urchin diversification. They were the first echinoids to achieve widespread biogeographical dispersal and achieved high levels of species richness compared to other clades of stem group echinoids. Following long-standing controversy regarding their phylogenetic affinities within the phylum Echinodermata, bothriocidaroids are now regarded as echinoids. The species- and genus-level phylogenetic relationships of the bothriocidaroids, are, however, less clearly known. We herein compile a database of bothriocidaroid occurrences, undertake detailed phylogenetic analyses of all named species within the Bothriocidaroida, and provide updated diagnoses for all species. Our phylogenetic analysis includes species belonging to three genera: Bothriocidaris, Unibothriocidaris and Neobothriocidaris. Furthermore, we describe a new species, Neobothriocidaris pentlandensis sp. nov., from the Silurian of Scotland and Sweden, using micro-computed tomography and scanning electron microscopy. This species has implications for understanding the diversity dynamics of bothriocidaroids before and after the Late Ordovician mass extinction and the palaeobiogeographical distribution of bothriocidaroids in the Silurian. Using Bayesian and parsimony-based phylogenetics, we demonstrate that Neobothriocidaris is a distinct clade, and depending upon the analytical technique, Unibothrioidaris and Bothriocidaris are also resolved as monophyletic. Unibothriocidaris appears to have become extinct in the Sandbian, while Neobothriocidaris and Bothriocidaris survived the Late Ordovician extinction events and lasted at least until the Ludlow. Furthermore, we performed statistical time-calibration of our phylogenetic trees and show that bothriocidaroids originated in the Dapingian or Darriwilian, diversifying in the Darriwilian to Sandbian and reaching peak diversity in the Sandbian and Katian. This Sandbian and Katian peak is similar to that observed in other echinoderm clades, as well as other animal groups during the Ordovician radiation, and may be linked to heightened sea level and expansive tropical shelves present in the Upper Ordovician, and to new inhabitable environments linked with Ordovician oxygenation. http://zoobank.org/urn:lsid:zoobank.org:pub:B55D0B22-D4AB-474A-80B9-D19BB43107F0","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"19 1","pages":"1395 - 1448"},"PeriodicalIF":2.6,"publicationDate":"2022-04-14","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"59822589","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"A re-assessment of the osteology and phylogenetic relationships of the enigmatic, large-headed reptile Sphodrosaurus pennsylvanicus (Late Triassic, Pennsylvania, USA) indicates archosauriform affinities","authors":"M. Ezcurra, H. Sues","doi":"10.1080/14772019.2022.2057820","DOIUrl":"https://doi.org/10.1080/14772019.2022.2057820","url":null,"abstract":"Most Triassic terrestrial diapsids belong to two clades, Lepidosauromorpha or (the more diverse) Archosauromorpha. Nevertheless, the phylogenetic relationships of some Triassic diapsids have remained uncertain for decades because of the lack of preservation of phylogenetically relevant anatomical regions or because of unusual combinations of features. One of these enigmatic forms is the small-sized Sphodrosaurus pennsylvanicus from the Upper Triassic Hammer Creek Formation of the Newark Supergroup in Pennsylvania (USA). It was first identified as a procolophonid parareptile, later as a probable rhynchosaur archosauromorph, and more recently as an indeterminate neodiapsid. Here we revise the anatomy of Sphodrosaurus pennsylvanicus in order to include it for the first time in a quantitative phylogenetic analysis, which is focused on Permo–Triassic neodiapsids. Sphodrosaurus pennsylvanicus is recovered in this analysis as a doswelliid proterochampsian within Archosauromorpha. As a result, this taxon is added to the list of doswelliids known from the Carnian–Norian of the eastern and south-western USA. Previous authors recognized that the most unusual feature of Sphodrosaurus pennsylvanicus is its proportionally very large skull. Phylogenetic generalized least squares regressions confirmed that Sphodrosaurus pennsylvanicus has a larger skull than the vast majority of Permo–Triassic diapsids. Optimization in the phylogeny of the skull width to presacral length ratio shows the most likely scenario is that the extremely broad skull of Sphodrosaurus pennsylvanicus is autapomorphic, but it is not unique among archosauromorphs, being paralleled by hyperodapedontine rhynchosaurs and the proterochampsian Proterochampsa barrionuevoi. Exploration of a morphospace of linear measurements shows that Sphodrosaurus pennsylvanicus shares strong similarities with the probably semi-aquatic Proterochampsa barrionuevoi, suggesting that the former species may have had a similar mode of life. A linear discriminant analysis of ungual functional categories found that the only preserved ungual of Sphodrosaurus pennsylvanicus was suitable for digging or some other way of substrate processing.","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"19 1","pages":"1643 - 1677"},"PeriodicalIF":2.6,"publicationDate":"2021-12-17","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"46613133","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}