Xinxin Ren, Shan Jiang, Xuri Wang, Guangzhao Peng, Y. Ye, Logan King, H. You
{"title":"Osteology of Dashanpusaurus dongi (Sauropoda: Macronaria) and new evolutionary evidence from Middle Jurassic Chinese sauropods","authors":"Xinxin Ren, Shan Jiang, Xuri Wang, Guangzhao Peng, Y. Ye, Logan King, H. You","doi":"10.1080/14772019.2022.2132886","DOIUrl":"https://doi.org/10.1080/14772019.2022.2132886","url":null,"abstract":"The Middle Jurassic lower Shaximiao Formation in Sichuan Province of south-western China has yielded a diverse terrestrial vertebrate fauna dominated by sauropod dinosaurs. However, many of these sauropods lack detailed descriptions or explicit phylogenetic diagnoses. Here, we present a comprehensive redescription of Dashanpusaurus dongi, a species of sauropod found only in the lower Shaximiao Formation. We define the revised autapomorphies of the species as follows: neural canals are sub-square in anterior dorsal vertebrae; the presence of a thin accessory lamina that contacts the prezygodiapophyseal and paradiapophyseal laminae of the middle dorsals, forming an angle of 75° to the horizontal; and four ridges on the anterodistal edge of the humerus. Often considered part of the epipophyseal-prezygapophyseal lamina, a strut invades the spinodiapophyseal fossa in the cervical and anterior dorsal vertebrae. Anatomical comparisons indicate that this feature was widespread among early-diverging Middle Jurassic eusauropod lineages. This comparative anatomical data provides an opportunity to revisit the phylogenetic position of Dashanpusaurus and the relationships of the neosauropod clade. Recovered as a macronarian, a better understanding of Dashanpusaurus dongi will allow for clarification of the origin, early evolution, and palaeogeographical distribution of neosauropods. This study also suggests that the diversity and dispersity of the neosauropod clade occurred much earlier than previously realized.","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":"1 - 72"},"PeriodicalIF":2.6,"publicationDate":"2022-11-18","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"48905904","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
G. Shi, Fabiany Herrera, P. Herendeen, E. G. Clark, P. Crane
{"title":"Silicified cupulate seed-bearing structures from the Early Cretaceous of eastern Inner Mongolia, China: rethinking the corystosperm concept","authors":"G. Shi, Fabiany Herrera, P. Herendeen, E. G. Clark, P. Crane","doi":"10.1080/14772019.2022.2133644","DOIUrl":"https://doi.org/10.1080/14772019.2022.2133644","url":null,"abstract":"The group referred to informally as the corystosperms, described initially based on compression fossils from the Triassic of Gondwana, have long been considered critical extinct plants for understanding seed plant phylogeny, the evolution of seed plant reproductive structures and the relationships of angiosperms. Here we describe a new genus and species of corystosperm seed-bearing structure, Jarudia zhoui gen. et sp. nov., based on abundant silicified material collected from the newly discovered chert in the Early Cretaceous Huolinhe Formation of eastern Inner Mongolia, north-eastern China. Jarudia zhoui is a lax seed cone consisting of a flexible central axis bearing deciduous, helically arranged, lateral seed-bearing units. Individual seed-bearing units consist of an elongate bract partially fused to an unbranched cupule stalk that bears a single, reflexed cupule apically. Each cupule is formed by the strongly reflexed cupule stalk and one median and two lateral flaps. The cupule stalk supplied by two vascular bundles and three unvascularized flaps partially enclose two three-angled seeds. Jarudia zhoui bears a striking resemblance to Doylea tetrahedrasperma from the Early Cretaceous of Canada and similar plants from the Early Cretaceous of Mongolia. There are also strong similarities with ovulate structures of Umkomasia from the Triassic of Gondwana in the structure and anatomy of individual cupules, their axial nature, and the architecture of the entire seed-bearing structure that has two orders of branching. New information from Jarudia zhoui, together with information on other corystosperm ovulate organs from the Northern Hemisphere, significantly expands our understanding of this key group of extinct plants, suggests that the cupules of the Early Cretaceous and Triassic corystosperms are homologous, and raises critical questions about the definition and phylogenetic circumscription of the corystosperms, including how Early Cretaceous and Triassic corystosperms are related to each other and to other groups of seed plants, including angiosperms.","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":"1 - 33"},"PeriodicalIF":2.6,"publicationDate":"2022-11-15","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"44607010","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"A new proterochampsid (Archosauriformes: Proterochampsia) from the Late Triassic of southern Brazil and the emergence of archosaurian hind limb traits","authors":"R. Müller, M. S. Garcia, André O Fonseca","doi":"10.1080/14772019.2022.2128913","DOIUrl":"https://doi.org/10.1080/14772019.2022.2128913","url":null,"abstract":"Characterized by an elongated snout, proterochampsids are carnivorous non-archosaur archosauriforms. The clade is endemic to South America and its fossil record extends from the early Carnian to the late Carnian/early Norian. Nesting close to Archosauria, it is a key clade for understanding the origin and evolution of archosaurian traits. Unfortunately, hind limb elements are usually poorly preserved for the group. Therefore, the hind limb anatomy of proterochampsids still lacks detailed descriptions. In the present study, we partially fill this gap with the description of a new proterochampsid represented by a complete and well-preserved hind limb. Stenoscelida aurantiacus gen. et sp. nov. was excavated from the late Carnian/early Norian (Late Triassic) beds of southern Brazil. A phylogenetic investigation recovers the new taxon as a non-rhadinosuchine proterochampsid. The species bears an unusual set of traits for the group, which provides clues on the evolutionary origins of some muscle attachment structures. For instance, the femur of Stenoscelida aurantiacus gen. et sp. nov. possesses an anterior trochanter and an anterolateral scar. So far, these features have not been reported in other non-archosaurian archosauriforms. Therefore, the new specimen indicates that some typical archosaurian features evolved earlier than previously thought. The taxon also carries additional uncommon features for proterochampsids, such as an iliofibularis tubercle on the anterior margin of the fibula and a vestigial phalanx in digit V. In sum, Stenoscelida aurantiacus has one of the best-preserved hind limbs within Proterochampsidae and sheds light on the polarization of important traits regarding the evolutionary context of Archosauria. http://zoobank.org/urn:lsid:zoobank.org:pub:C78B7CE3-AB9B-4543-833E-B2A3FEA957D9","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":"1 - 19"},"PeriodicalIF":2.6,"publicationDate":"2022-11-03","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"45706780","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
M. Pulsipher, Evan P. Anderson, Lauren S. Wright, J. Kluessendorf, D. Mikulic, J. Schiffbauer
{"title":"Description of Acheronauta gen. nov., a possible mandibulate from the Silurian Waukesha Lagerstätte, Wisconsin, USA","authors":"M. Pulsipher, Evan P. Anderson, Lauren S. Wright, J. Kluessendorf, D. Mikulic, J. Schiffbauer","doi":"10.1080/14772019.2022.2109216","DOIUrl":"https://doi.org/10.1080/14772019.2022.2109216","url":null,"abstract":"The Silurian Waukesha Lagerstätte of Wisconsin, USA, hosts a fauna of distinctive soft-bodied organisms, including a range of graptolites, worms, trilobites and uncommon arthropods, but otherwise lacks most of the shelly fauna typical of contemporaneous deposits. While several species have been described, much of the Waukesha biota remains understudied. Here, we examine numerous specimens of a previously unnamed and undescribed vermiform arthropod that bears a cephalic carapace, a suite of specialized head appendages, a long, segmented trunk, and a battery of small swimming appendages. The taxonomic and phylogenetic evaluations presented herein establish this taxon as a novel genus and species, Acheronauta stimulapis gen. et sp. nov., plausibly positioned within the basal mandibulates. Our phylogenetic analyses suggest that a new arthropod clade may be warranted, comprising Acheronauta gen. nov., Captopodus, and the thylacocephalans. https://zoobank.org/urn:lsid:zoobank.org:pub:B572831B-1A8A-4E4E-9258-34D91AA22A74nk","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":"1 - 24"},"PeriodicalIF":2.6,"publicationDate":"2022-10-10","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"45996007","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Phylogeny of the amphibamiform temnospondyls: the relationship of taxa known by adults, larvae and neotenes","authors":"R. Schoch","doi":"10.1080/14772019.2022.2113831","DOIUrl":"https://doi.org/10.1080/14772019.2022.2113831","url":null,"abstract":"The Amphibamiformes, small temnospondyls from late Palaeozoic rocks, have been increasingly considered as the stem-group of some or all extant amphibians (Lissamphibia). Their relationships have become intensely studied after the discovery of new taxa and the revision of poorly known ones, but understanding is hampered by the fact that amphibamiforms fall into two very distinct groups: (a) terrestrial taxa known by adults only (Amphibamidae, Micropholidae) and (b) aquatic taxa known by larvae and neotenic adults (Branchiosauridae). The Branchiosauridae were recognized as a clade supported by a suite of larval synapomorphies, but the unknown larval morphology of the terrestrial clades leaves the question open to whether some branchiosaurid apomorphies might have been more widely distributed. Recently, some branchiosaurid specimens from Nýřany (Czech Republic) were proposed to be larvae of olsoniforms and amphibamids, and together with a revision of ancient branchiosaurid-like taxa from the United States and France prompt a re-analysis of this issue. In a phylogenetic analysis of 48 taxa and 113 characters, the Branchiosauridae were found to be monophyletic and nest in an unresolved trichotomy with Amphibamidae and the putative relatives of Lissamphibia. The present analysis finds the hypothesized olsoniform larvae from Nýřany to be adults of Branchiosaurus salamandroides, whereas some of the larvae from Mazon Creek (Illinois, USA) are not branchiosaurids, but immature specimens of the amphibamid Amphibamus grandiceps. Within branchiosaurids, four clades have been identified: (a) the Carboniferous Anthracobamus clade (Anthracobamus, Montceaubatrachus), (b) the mostly Carboniferous Branchiosaurinae (Branchiosaurus, Milnererpeton) and two largely Permian clades, the newly defined Apateoninae (genus Apateon) and Leptorophinae (Melanerpeton, Leptorophus, Schoenfelderpeton) clade. Branchiosaurid evolution involved a trend towards an enhanced paedomorphosis, a modification of branchial teeth, and successive reduction of bone in the scales and pectoral girdle.","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":"1 - 30"},"PeriodicalIF":2.6,"publicationDate":"2022-10-05","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"46847616","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Careful amendment of morphological data sets improves phylogenetic frameworks: re-evaluating placement of the fossil Amiskwia sagittiformis","authors":"Nicolas Bekkouche, L. Gąsiorowski","doi":"10.1080/14772019.2022.2109217","DOIUrl":"https://doi.org/10.1080/14772019.2022.2109217","url":null,"abstract":"The Cambrian fossil Amiskwia sagittiformis has puzzled palaeontologists for more than a century, but recent re-investigation of its morphology suggested a close relationship with the bilaterian clade Gnathifera, comprising Rotifera, Gnathostomulida and Micrognathozoa. Since Amiskwia has already been considered closely related to Chaetognatha, this new interpretation of its morphology supports recent molecular and developmental studies finding a close phylogenetic relationship between Chaetognatha and Gnathifera. The recent re-description of the jaw apparatus of Amiskwia with the first phylogenetic analysis by Vinther & Parry (2019) found a rather surprising topology with Amiskwia and Chaetognatha being sister groups and nested inside Gnathifera. Furthermore, a subsequent paper from Caron & Cheung (2019) re-described the jaws of Amiskwia as very similar to the configuration found in Gnathostomulida, but did not include any phylogenetic analysis. Here we test the topology of Vinther & Parry with various parsimony and Bayesian analyses, taking into account the new description of Caron & Cheung with careful re-amendment of the matrix of Vinther & Parry. According to our results and the recent findings on the molecular phylogeny of their extant members, we suggest a new systematization of these taxa. We recovered Amiskwia as a stem-group chaetognath within a clade Cucullophora nov., rejecting the Caron & Cheung hypothesis of its close affinity to Gnathostomulida, and found a more consensual topology with monophyletic Gnathifera sister group to Cucullophora, all together forming the clade Chaetognathifera. Furthermore, Rotifera + Micrognathozoa form a clade named Gynognathifera nov. We discuss the characters supporting each clade and the reasons that account for the topology found by Vinther & Parry. Finally, we hope that this carefully amended matrix focused on Gnathifera, Amiskwia and Chaetognatha, together with a formal classification and robust phylogeny, will be of use for future studies on the palaeontology and morphology of these clades.","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":"1 - 14"},"PeriodicalIF":2.6,"publicationDate":"2022-10-05","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"47802911","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Re-description and phylogenetic relationships of †Protosyngnathus sumatrensis (Teleostei: Syngnathoidei), a freshwater pipefish from the Eocene of Sumatra, Indonesia","authors":"Alison M. Murray","doi":"10.1080/14772019.2022.2113832","DOIUrl":"https://doi.org/10.1080/14772019.2022.2113832","url":null,"abstract":"†Protosyngnathus sumatrensis is known from six specimens from Eocene lacustrine deposits of the Sangkarawang Formation of Sumatra, Indonesia. The fish was named almost 150 years ago, but has never been well described or figured. Probably because of this, it has not been included in phylogenies of extant pipefishes and seahorses (Syngnathiformes). A detailed description is provided, along with a new phylogenetic analysis of morphological characters that allows the Eocene fish to be confidently placed with the extant families Syngnathidae (pipefishes and seahorses) and Solenostomidae (ghost seahorses) along with the Paleocene marine fossil †Eekaulostomus cuevasae from Mexico. The vast majority of these fishes are marine inhabitants, with a few syngnathids entering brackish or freshwater habitats. †Protosyngnathus documents the occurrence of syngnathiform fishes in a lacustrine environment, indicating that this lineage invaded and became adapted to the freshwater environment early in the evolution of this group.","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":"1 - 16"},"PeriodicalIF":2.6,"publicationDate":"2022-09-30","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"46856239","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"The first evidence of Heosminthus from North America and the phylogenetics of Sminthidae (Mammalia, Rodentia, Dipodoidea): biogeographical implications","authors":"J. Calede, Y. T. Tse, K. D. Cairns","doi":"10.1080/14772019.2022.2111232","DOIUrl":"https://doi.org/10.1080/14772019.2022.2111232","url":null,"abstract":"The Eurasian fossil record is rich with the remains of sminthid rodents, a clade today represented by a single genus, Sicista. In contrast, the North American record of the family is sparse and exclusively composed of species of Plesiosminthus, an animal bearing a grooved incisor, during the Oligocene. This fossil record is the product of dispersal events from Eurasia. We here present the first occurrence of a smooth-incisor sminthid from North America. The new species, Heosminthus teredens from the Arikareean-aged Cabbage Patch beds of Montana, represents a rare occurrence of cranial material showing the clear association between incisors and cheek teeth, and the first postcranial remains for fossil sminthids. Our phylogenetic analysis, expanding on prior work, shows that Heosminthus teredens is most closely related to Heosminthus borrae, from Mongolia. In combination with published radioisotopic dating, our phylogenetic framework of sminthids provides evidence for an additional dispersal of sminthid rodents from Eurasia to North America 30 million years ago via Beringia. Our morphometric analysis of the tibia and metatarsal supports a terrestrial ecology for the new species; it is possible the animal had limited scansorial and/or saltatorial abilities. The large sample size of teeth available for the new taxon enables an analysis of morphological variation and indicates a high degree of polymorphism as well as a large size variation, even when controlling for stratigraphical and geographical range. https://zoobank.org/urn:lsid:zoobank.org:pub:B7D4245C-992F-4956-96AD-10D7CFA8E5C8","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":"1 - 21"},"PeriodicalIF":2.6,"publicationDate":"2022-09-22","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"41811530","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
M. A. Cerroni, M. Baiano, J. Canale, F. Agnolín, A. Otero, F. Novas
{"title":"Appendicular osteology of Skorpiovenator bustingorryi (Theropoda, Abelisauridae) with comments on phylogenetic features of abelisaurids","authors":"M. A. Cerroni, M. Baiano, J. Canale, F. Agnolín, A. Otero, F. Novas","doi":"10.1080/14772019.2022.2093661","DOIUrl":"https://doi.org/10.1080/14772019.2022.2093661","url":null,"abstract":"Skorpiovenator bustingorryi is a derived abelisaurid theropod represented by a fairly complete skeleton from the Late Cretaceous sedimentary beds of north-western Patagonia. Although some features were described in the original paper, mainly related to the skull, the appendicular anatomy remains undescribed. The aim of the present contribution is to provide a detailed description and analysis of the available appendicular bones, including comparisons with other ceratosaurian theropods close to Skorpiovenator. In this way, new autapomorphies emerged to further distinguish Skorpiovenator from its relatives. Furthermore, a comprehensive phylogenetic analysis was performed and several characteristics of the hind limb, in particular some of the autopodium, resulted in the identification of new apomorphic traits for Ceratosauria and Abelisauridae. These features might prove to be useful for future phylogenetic analyses and may help to resolve the still confusing and debated internal relationships of abelisaurid theropods.","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":"1 - 32"},"PeriodicalIF":2.6,"publicationDate":"2022-09-22","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"47879157","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"A nearly complete skeleton of a new eusphenodontian from the Upper Jurassic Morrison Formation, Wyoming, USA, provides insight into the evolution and diversity of Rhynchocephalia (Reptilia: Lepidosauria)","authors":"D. Demar, Marc E. H. Jones, M. Carrano","doi":"10.1080/14772019.2022.2093139","DOIUrl":"https://doi.org/10.1080/14772019.2022.2093139","url":null,"abstract":"We describe a new, small-bodied rhynchocephalian reptile, Opisthiamimus gregori gen. et sp. nov., from the Upper Jurassic Morrison Formation of Wyoming, USA. Whereas many fossil rhynchocephalians are based on isolated incomplete jaws, the holotype of O. gregori includes most of the skull and postcranium and therefore represents one of the most complete specimens of Rhynchocephalia known from North America. We used micro-computed tomography to examine its skeletal anatomy in detail and to develop a three-dimensional reconstruction of the skull. The skull of O. gregori is similar to that of several non-neosphenodontian rhynchocephalians such as Planocephalosaurus (e.g. large orbits) and Clevosaurus (e.g. parietal parasagittal crests) yet exhibits a suite of other features related to the proal shearing mechanism that becomes increasingly elaborated among more phylogenetically nested taxa such as Sphenodon (e.g. lateral palatine tooth row parallels maxillary tooth row along its entire length, pyramidal dentary teeth with mesial shearing crests). The postcranial skeleton of O. gregori exhibits characteristics typical of a terrestrial rhynchocephalian. Our phylogenetic analyses use a substantially updated data set of 118 characters and 46 taxa, and both maximum parsimony and Bayesian frameworks. Results place O. gregori inside Eusphenodontia but outside Neosphenodontia, and therefore in a key position for contributing to character polarity for more deeply nested clades such as Clevosauridae, Sphenodontidae and Pleurosauridae. We also erect Leptorhynchia taxon nov., composed primarily of aquatically adapted taxa (e.g. Pleurosaurus, Sapheosaurus), which is supported by both cranial and postcranial characters. Because O. gregori is not particularly closely related to the other named Morrison rhynchocephalians (e.g. Opisthias rarus), it increases both the alpha and beta taxonomic diversities within the formation. Similarly, major differences in body size and inferred diet of the Morrison taxa imply considerable concomitant palaeoecological diversity just prior to a major global decline in rhynchocephalian diversity around the close of the Jurassic. http://zoobank.org/urn:lsid:zoobank.org:pub:888E055B-8AC1-4BD0-A37C-8CB192F79673","PeriodicalId":50028,"journal":{"name":"Journal of Systematic Palaeontology","volume":"20 1","pages":"1 - 64"},"PeriodicalIF":2.6,"publicationDate":"2022-09-15","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"41879119","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}