Qi Zhang , Ning Jia , Yizheng Li , Jianxun Wang , Alexander P. Rasnitsyn
{"title":"New genus of falsiformicid wasps (Hymenoptera, Vespoidea: Falsiformicidae) with brachypterous male from middle Cretaceous Kachin amber in Myanmar","authors":"Qi Zhang , Ning Jia , Yizheng Li , Jianxun Wang , Alexander P. Rasnitsyn","doi":"10.1016/j.cretres.2024.106057","DOIUrl":"10.1016/j.cretres.2024.106057","url":null,"abstract":"<div><div>Falsiformicidae is a poorly known extinct family of aculeate wasps. It is known only as inclusions in various Cretaceous ambers. Brachypterous male <em>Falsiformix pedestris</em> Zhang and Rasnitsyn gen. et sp. nov. is described in the family Falsiformicidae based on a unique fossil from the middle Cretaceous Kachin amber. Occurrence and meaning of the male flightlessness in Hymenoptera is discussed. A combination of the flightless males with flying females occurs when mating takes place shortly after the eclosion, before adults disperse from confined space where they developed. An equally rare even if less exotic strategy with males and females both being flightless depends on the female pedestrian habits rather than on the pre-dispersal mating. Wide spatial (Angara Land, Laurussia Land and Gondwana Land) and narrow temporal distribution (Cenomanian and Turonian ages only) indicate Falsiformicidae as a possibly important stratigraphical indicator.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"168 ","pages":"Article 106057"},"PeriodicalIF":1.9,"publicationDate":"2024-11-26","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"143181848","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Aijing Li , Baoxia Du , Jing Zhang , Jing Peng , Yiqiao Fu , Jingjing Cai , Mingyang Wei , Peihong Jin
{"title":"A new species Torreya with the seed-bearing structure from the Lower Cretaceous of northwestern China and its evolutionary significance","authors":"Aijing Li , Baoxia Du , Jing Zhang , Jing Peng , Yiqiao Fu , Jingjing Cai , Mingyang Wei , Peihong Jin","doi":"10.1016/j.cretres.2024.106056","DOIUrl":"10.1016/j.cretres.2024.106056","url":null,"abstract":"<div><div>Taxaceae is a special member of the Mesozoic flora mainly found in the Northern Hemisphere. However, Taxaceae fossils did not show abundance in the Mesozoic, particularly lacking twigs with information on reproductive organs. The new species <em>Torreya jiuquanensis</em> sp. nov. from the Lower Cretaceous of the Jiuquan Basin, Gansu, NW China provides valuable insights in seed-bearing structures of <em>Torreya</em>. Firstly, we present a comprehensive morphological description of <em>Torreya jiuquanensis</em> and it is distinguished from analogous fossils by more slender leaves and the obovate ovule. Secondly, phylogenetic analysis was utilized to determine that the position of <em>Torreya jiuquanensis</em> is located in the same clade as <em>Torreya taxifolia</em>. Meanwhile, the remarkable traits of <em>Torreya jiuquanensis</em> indicated that <em>Torreya</em> had demonstrated morphological stasis at least since the Early Cretaceous. Finally, through the examination of other genera within Taxaceae, it proposed that morphological stasis is not only observed in <em>Torreya</em> but also prevalent across nearly all genera of the Taxaceae, which are characterized by an evolutionary hysteresis. Moreover, a reinvestigation of the global fossil records and paleogeographic distribution of <em>Torreya</em> have revealed that <em>Torreya</em> was primarily restricted to the Northern Hemisphere in the Mesozoic, and multiple climate fluctuations were key drivers behind shifts in <em>Torreya</em> diversity and range migration.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"168 ","pages":"Article 106056"},"PeriodicalIF":1.9,"publicationDate":"2024-11-21","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"143181578","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
{"title":"Two new species of the extinct subfamily Alitrepaninae (Coleoptera: Bostrichidae) from the Upper Cretaceous (Cenomanian) Kachin amber in northern Myanmar","authors":"Huan Wang, Yuan Peng, Qianhui Lin, Ruiqing Tao, Zirui Zhang, Shuo Wang","doi":"10.1016/j.cretres.2024.106051","DOIUrl":"10.1016/j.cretres.2024.106051","url":null,"abstract":"<div><div>Bostrichidae is commonly referred to as auger beetles. Of them, the subfamily Alitrepaninae is the only extinct subfamily of the family Bostirichidae. Here, based on detailed characterization analysis and comparisons with other species of the Alitrepaninae subfamily, we identified two new species of the genus <em>Poinarinius</em> in well-preserved Upper Cretaceous (Cenomanian) amber from northern Myanmar, along with a key to <em>Poinarinius</em> genus. Our new species can be clearly distinguished through the feature of head visible from above, dorsal plate of the pronotum, elytra performance and length of each tarsomere. Our study deepens insight into the adaptive evolution of the subfamily Alitrepaninae regarding feeding habits and resistance, as well as discussed the ecological habits of this taxon in Cretaceous. In summary, our findings will contribute to the understanding of the diversity and evolutionary origin of Bostirichidae.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"167 ","pages":"Article 106051"},"PeriodicalIF":1.9,"publicationDate":"2024-11-20","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"142747482","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Ángela D. Buscalioni , Raman Umamaheswaran , Alba Sánchez-García , Ana Isabel López-Archilla , Jaime Joaquim Dias , Ismar de Souza Carvalho , Candela Blanco-Moreno , Jesús Marugán-Lobón
{"title":"Exceptional preservation in mayfly nymphs (Insecta: Ephemeroptera) from the Early Cretaceous of the Las Hoyas fossil site","authors":"Ángela D. Buscalioni , Raman Umamaheswaran , Alba Sánchez-García , Ana Isabel López-Archilla , Jaime Joaquim Dias , Ismar de Souza Carvalho , Candela Blanco-Moreno , Jesús Marugán-Lobón","doi":"10.1016/j.cretres.2024.106052","DOIUrl":"10.1016/j.cretres.2024.106052","url":null,"abstract":"<div><div>The present study focuses on a group of macroinvertebrates that have played an important role in past and present lentic ecosystems: the Ephemeroptera. The study analyses the diversity, relative abundance, taphonomic alterations and fossilisation patterns of the Las Hoyas nymphs. The objective is to investigate the factors that may contribute to the presence of soft tissues in fossils with poor corporeal preservation. In accordance with this, only three morphotypes of the Leptophlebiidae family and one of the Euthyplociidae family have been characterised. However, the ultrastructural details of digestive and respiratory tissues have been documented, including the digestive canal, the shape of the gills, tracheoles, chloride cells, and importantly, in situ bacteria in cytokinetic stages located at the hindgut. The nymphs were fossilised in a variety of ways, including carbonaceous films, mineralised casts, and impressions. The nymphs that have been fossilised as composite moulds (Type III) and carbon films (Type II) tend to be larger in size. However, their bodies are rather well-preserved and show less soft tissue remains than those of the nymphs that have been preserved as impressions (Type I) and mineralised phosphorus casts (Type IV). These latter nymphs are smaller in size, and their bodies are poorly preserved. However, they show a greater variety of soft tissue details. This bias suggests an interesting link to environmental factors differentiating watered and flooding from microbial mat binding conditions.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"167 ","pages":"Article 106052"},"PeriodicalIF":1.9,"publicationDate":"2024-11-19","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"142747481","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"OA","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Yan Zhang , Yun-fei Xue , Chuan-biao Wan , Wen-chun Ge
{"title":"A new palynological assemblage of the Nantun Formation in the Huhehu Sag, Hailar Basin, NE China: Implications for paleoenvironments","authors":"Yan Zhang , Yun-fei Xue , Chuan-biao Wan , Wen-chun Ge","doi":"10.1016/j.cretres.2024.106054","DOIUrl":"10.1016/j.cretres.2024.106054","url":null,"abstract":"<div><div>The Cretaceous period, recognized as one of the warmest in Earth's geological timeline, is crucial for understanding paleoclimatic conditions. Palynological analyses, focusing on spores and pollen, play a vital role in reconstructing paleoenvironmental. This study emphasizes the relationship between climate and plant distribution through detailed palynological analyses. It reports the first discovery of abundant and diverse palynomorphs from the lower Nantun Fm in the Hailar Basin, Inner Mongolia, China. The identified palynological assemblage, termed the <em>Protoconiferus flavus</em>—<em>Pinuspollenites</em> sp.—<em>Quantouendinium</em> sp. (PPQ) assemblage, provides significant insights into the Early Cretaceous flora of northeastern China. The PPQ assemblage can be chronologically attributed to the Hauterivian–early Barremian interval, consistent with the LA–ICP–MS zircon U–Pb isotope ages of the Nantun Fm volcanic rocks. Paleovegetation reconstruction reveals a prevailing dominance of coniferous vegetation in the region represented by the PPQ assemblage, indicative of a characteristic of mid-to high-latitude mountain coniferous forest landscape. The climatic was mainly humid or semi-humid. This conclusion is supported by the abundance of freshwater dinocyst and green algae fossils, suggesting that sedimentation occurred in a shallow, freshwater hydrostatic environment with an estimated water temperature of approximately 20–25 °C. The environment was also nutrient-rich, which facilitated the proliferation of green algae such as <em>Scenedesmus</em>, <em>Pediastrum</em>, and <em>Tetraedron</em>. Identifying this assemblage is important for determining the age of the lower Nantun Fm, reconstructing paleovegetation and paleoclimate, and analyzing the lacustrine water conditions during the sedimentary period in the Hailar Basin.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"167 ","pages":"Article 106054"},"PeriodicalIF":1.9,"publicationDate":"2024-11-19","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"142747480","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
P.E. Santamarina , V.S. Perez Loinaze , M.S. González Estebenet , M.V. Guler , C. Discenza , S. Lizzoli , A.N. Varela
{"title":"Palynology from the Cenomanian Puesto El Moro Formation (Santa Cruz Province, Argentina)","authors":"P.E. Santamarina , V.S. Perez Loinaze , M.S. González Estebenet , M.V. Guler , C. Discenza , S. Lizzoli , A.N. Varela","doi":"10.1016/j.cretres.2024.106055","DOIUrl":"10.1016/j.cretres.2024.106055","url":null,"abstract":"<div><div>During the Late Cretaceous and the Paleocene, terrestrial floras witnessed the so-called “Angiosperm Terrestrial Revolution”, when several modern clades of angiosperms originated and diversified. This process is scarcely studied in the Southern Hemisphere. Also, during the Cretaceous several carbon cycle alterations were registered, producing climatic modifications, and affecting both continental and marine biota. We present the palynological composition of the Puesto El Moro Formation, from the Cenomanian of Santa Cruz Province, Argentina. The sequence shows a progressive continentalization from coastal to fluvial sediments, representing an opportunity to understand how terrestrial floras evolved during this period in the Southern Hemisphere and how global climatic events affected them. Fifteen fertile palynological samples were studied. 142 species were recovered, with 23 bryophytes and lycophytes, 64 ferns, 34 gymnosperms, and 13 angiosperms. Dinoflagellate cysts are moderately abundant in the basal levels, but have poor preservation. The age of the unit was previously established as Cenomanian based on stratigraphic relations, and the palynostratigraphy supports this conclusion. Counts show that almost all the assemblages are dominated by gymnosperms (primarily <em>Classopollis</em>), followed by ferns (primarily <em>Cyathidites</em>). Bryophytes, lycophytes and angiosperms represent a minor proportion. Richness analysis shows that, on average, it tends to increase, with a great rise in the uppermost levels, pointing out the possible location of the Mid-Cenomanian Event. The Puesto El Moro palynoflora is closely related with the ones from the <em>Classopollis</em>-<em>Cyclusphaera</em> palynological sub-province. The analysis shows a latitudinal arrangement of the palynofloras, differentiated between them by their most abundant gymnosperm pollen grain.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"168 ","pages":"Article 106055"},"PeriodicalIF":1.9,"publicationDate":"2024-11-19","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"143180243","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Martina Kočová Veselská , Mikhail Rogov , Rok Gašparič , Matúš Hyžný , Alexey P. Ippolitov , Javier Luque , Martin Košťák
{"title":"A unique insight into the growth of necrocarcinoid crabs (Malacostraca: Decapoda: Brachyura): Evidence from the Upper Cretaceous of Siberia (Russia), with a description of a new species","authors":"Martina Kočová Veselská , Mikhail Rogov , Rok Gašparič , Matúš Hyžný , Alexey P. Ippolitov , Javier Luque , Martin Košťák","doi":"10.1016/j.cretres.2024.106053","DOIUrl":"10.1016/j.cretres.2024.106053","url":null,"abstract":"<div><div>The Upper Cretaceous (Cenomanian to Coniacian, 100.5–86.3 Ma) strata of the northern Krasnoyarsk Krai (Russia) yielded new fossil brachyuran crab material, based on which a new species, <em>Necrocarcinus sibiricus</em>, is described. Although the specimens come from settings of slightly different ages, all of them display dorsal carapace characteristics consistent with variation attributable to ontogenetic changes related to growth, for instance, the narrowing of the orbitofrontal margin, and a continuous smoothing and widening of the carapace. Interestingly, due to these ontogenetic changes, the largest of the studied specimens exhibits a morphology close to that previously described in <em>Camarocarcinus</em>, a related necrocarcinoid crab genus known from Paleocene strata. The newly described species shows that the dorsal carapace morphology typical for <em>Camarocarcinus</em> can be observed in at least some fully grown necrocarcinids, prompting a re-evaluation of the relationship between Necrocarcinidae and Camarocarcinidae. We propose that camarocarcinids may be derived from within Necrocarcinidae, with the particular morphology of <em>Camarocarcinus</em> resulting from heterochronous developmental processes, i.e., peramorphosis.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"168 ","pages":"Article 106053"},"PeriodicalIF":1.9,"publicationDate":"2024-11-19","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"143180242","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Xuanyu Zhou , Naoki Ikegami , Rodrigo V. Pêgas , Toru Yoshinaga , Takahiro Sato , Toshifumi Mukunoki , Jun Otani , Yoshitsugu Kobayashi
{"title":"Reassessment of an azhdarchid pterosaur specimen from the Mifune Group, Upper Cretaceous of Japan","authors":"Xuanyu Zhou , Naoki Ikegami , Rodrigo V. Pêgas , Toru Yoshinaga , Takahiro Sato , Toshifumi Mukunoki , Jun Otani , Yoshitsugu Kobayashi","doi":"10.1016/j.cretres.2024.106046","DOIUrl":"10.1016/j.cretres.2024.106046","url":null,"abstract":"<div><div>The Japanese pterosaur record is relatively scarce and represented by a limited number of fragmentary specimens from Cretaceous deposits, including a partial cervical vertebra of an azhdarchid from the “Upper Formation” (Turonian-Coniacian) of the Mifune Group in Kumamoto Prefecture, on the island of Kyushu, Japan. Here, we redescribe this notable Japanese pterosaur specimen and test its phylogenetic position. We interpret it as a sixth cervical vertebra and identify diagnostic features that enable its recognition as a new taxon, <em>Nipponopterus mifunensis</em> gen. et sp. nov. It is noteworthy that this represents the first nominal species of pterosaur from Japan and that this new taxon shows numerous quetzalcoatline features, being strikingly similar to the unnamed Burkhant azhdarchid from the Turonian–Coniacian of Mongolia. Our phylogenetic analysis places <em>Nipponopterus</em> as a sister taxon to the Burkhant azhdarchid and nested within the clade of Quetzalcoatlinae.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"167 ","pages":"Article 106046"},"PeriodicalIF":1.9,"publicationDate":"2024-11-16","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"142704623","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
E. Aldrin Juárez-Aguilar , Francisco Sánchez-Beristain , Juan Pablo Bernal , Ofelia Morton-Bermea , Pedro García-Barrera
{"title":"Palaeoenvironmental inferences of a Mexican Konservat-Lagerstätte (Tlayúa Quarry; Lower Cretaceous) based on the geochemistry of rare earth elements","authors":"E. Aldrin Juárez-Aguilar , Francisco Sánchez-Beristain , Juan Pablo Bernal , Ofelia Morton-Bermea , Pedro García-Barrera","doi":"10.1016/j.cretres.2024.106045","DOIUrl":"10.1016/j.cretres.2024.106045","url":null,"abstract":"<div><div>The Tlayúa Quarry is considered one of the most important Cretaceous outcrops in Mexico. Its well-preserved palaeobiota has allowed this outcrop to be considered a <em>Konservat-Lagerstätte</em>. Geochemical proxies such as major, minor and trace elements were evaluated from selected Zones of the Tlayúa Quarry, in order to infer some features of the depositional environment in which Tlayúa limestones originated. Aluminum, Thorium and Zirconium concentrations in addition to a low Phosphorous content evidence little terrigenous contamination, as well as a phosphate-free environment. Additionally, proxies such as Fe vs ΣREE, Mn vs ΣREE, Al vs ΣREE, Fe vs Nd<sub>SN</sub>/Yb<sub>SN</sub>, Al vs Nd<sub>SN</sub>/Yb<sub>SN</sub>, and Ce/Ce∗ vs Eu/Eu∗ show low correlation values (r<sup>2</sup> less than 0.25 in all cases with n = 128 and p < 0.05), thus suggesting a negligible influence of diagenetic effects or terrigenous contamination. These conditions are necessary for carrying out reliable palaeoenvironmental interpretations. Rare Earth Elements + Yttrium (REE+Y) concentrations were normalised to the Post-Archean Australian Shale standard to evaluate REE+Y patterns and anomalies. Following traits were identified: a slightly enrichment of heavy REE relative to light REE and moderate REE; positive La/La<sub>SN</sub>, Gd/Gd<sub>SN</sub> and Y/Y<sub>SN</sub> anomalies; negative/positive Ce/Ce<sub>SN</sub> anomalies; superchondritic and chondritic Y/Ho ratios. Here, we described that most of our limestone samples show the influence of marine waters and well oxidising conditions. However, freshwater signals that have been reported for some regions of the Tlayúa Quarry were also verified. Geochemical data are in agreement with the depositional environment evidenced by microfacies.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"167 ","pages":"Article 106045"},"PeriodicalIF":1.9,"publicationDate":"2024-11-15","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"142704722","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Marouf Abdelaty Mohamed Abdelhamid , Osman Abdelghany , Mahmoud Abu Saima , Anhar Asan
{"title":"Selected regular echinoids (Echinoidea) from the upper Campanian–Maastrichtian along the western borders of the Northern Oman Mountains, with description of a new species","authors":"Marouf Abdelaty Mohamed Abdelhamid , Osman Abdelghany , Mahmoud Abu Saima , Anhar Asan","doi":"10.1016/j.cretres.2024.106037","DOIUrl":"10.1016/j.cretres.2024.106037","url":null,"abstract":"<div><div>Thirteen species of regular echinoid, in ten genera, are recorded from the upper Campanian–Maastrichtian Qahlah and Simsima formations of two sections located along the western borders of the Northern Oman Mountains. Twelve of these are from the Simsima Formation in the Jebel El Rawdah section, four from the Qahlah and Simsima formations in the Jebel Huwayyah section. One new species, <em>Hattopsis muradi</em>, is described from the Jebel El Rawdah section. The status of the genus <em>Pseudorthopsis</em> is confirmed; differences between it and <em>Orthopsis</em> are discussed and <em>Orthopsis sanfilippoi</em> is transferred to <em>Pseudorthopsis</em>, here being recorded for the first time from the Arabian Gulf region. Revision of three species referred to <em>Hattopsis</em> and nine species ascribed to <em>Protechinus</em> (=<em>Noetlingia</em>, <em>Noetlingaster</em>) has elucidated the relationship between these two genera and affirmed the distinctiveness of <em>Hattopsis</em>. <em>Hattopsis sphericus</em> and <em>Echinotiara perebaskinei</em> are recorded for the first time from Jebel Huwayyah. All species identified are compared with other related taxa, and their stratigraphical ranges, affinities, palaeobiogeography and palaeoecology discussed.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"167 ","pages":"Article 106037"},"PeriodicalIF":1.9,"publicationDate":"2024-11-09","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"142704622","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}