Rivista Italiana Di Paleontologia E Stratigrafia最新文献

{"title":"OLENEKIAN TO EARLY LADINIAN STRATIGRAPHY OF THE WESTERN PART OF THE AGHDARBAND WINDOW (KOPEH-DAG, NE IRAN)","authors":"M. Balini, A. Nicora, S. Zanchetta, A. Zanchi, R. Marchesi, I. Vuolo, Maryam Hosseiniyoon, M. Norouzi, S. Soleimani","doi":"10.13130/2039-4942/11446","DOIUrl":"https://doi.org/10.13130/2039-4942/11446","url":null,"abstract":"The structural setting and the stratigraphy of the Early to Middle Triassic sedimentary succession exposed in the western part of the Aghdarband window (Kopeh Dag, NE Iran) is described. Six stratigraphic sections in the Sefid-Kuh Limestone, Nazar-Kardeh Formation and Sina Formation have been studied in the tectonic units 1a and 2. The lithostratigraphy is revised, with bio-chronostratigraphic constrain provided by conodonts and ammonoids. The new Olenekian ammonoid genus Megatirolitesis erected. It is based on species thus far known only in Mangyshlak (West Kazakhstan) but it is occurs also in the Sefid-Kuh Limestone.The evolution of the Lower Triassic carbonate ramp of the Sefid-Kuh Limestone, persisted in the Middle Anisian, with a three-stage development (Sefid-Kuh 1, 2 and 3 members) separated by drowning and onset of siliciclastics. The last stage is in part coeval with the Middle Anisian basinal Nazarkardeh Formation.The unconformity-bounded, three-stage development of the carbonate ramp documents that in the Aghdarband Basin the tectonic control over sedimentation started already in the Olenekian, since the onset of the marine transgression. The transgression of the Ladinian Sina Formation sealed a complex morphology resulting from the uplift and erosion of the Middle Anisian units. A new paleogeographic position along the southern Laurasia margin is propsed for the Triassic Aghdarband Basin. Based on the paleobiogeographic affinity of the Olenekian ammonoid occurences, we suggest that the Aghdarband Basin was located in a back-arc position in close connection with Mangyshlak (West Kazakhstan) and Tuarkyr (Turkmenistan), passing northwestward to a large epicontinental basin extending to the Donbass area. At least during the Olenekian the Aghdarband Basin had no direct connection with the Nakhlak Basin, which was proably located in a different intra-arc or more probably fore-arc region with respect to the Palaeotethys subduction-related Triassic arc.","PeriodicalId":54451,"journal":{"name":"Rivista Italiana Di Paleontologia E Stratigrafia","volume":"125 1","pages":"283-315"},"PeriodicalIF":2.3,"publicationDate":"2019-03-25","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"46223336","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"Adria as promontory of Africa and its conceptual role in the tethys twist and pangea b to pangea a transformation in the permian","authors":"G. Muttoni, D. Kent","doi":"10.13130/2039-4942/11437","DOIUrl":"https://doi.org/10.13130/2039-4942/11437","url":null,"abstract":"It has been almost 60 years since the first results from the Early Permian Bolzano Quartz Porphyries from the Trento Plateau of northern Italy (Southern Alps) showed paleomagnetic inclinations steeper than inclinations from broadly coeval units from central Europe. This experimental discrepancy, confirmed ever since at varying levels of magnitude and certitude, implied that northern Italy had paleolatitudes too northerly relative to Europe to be considered part of the European continent. On the other hand, it became progressively more apparent that paleomagnetic data from northern Italy were more compatible with data from Africa than with data from Europe, and this observation revived and complemented Argand’s original concept of Adria as a promontory of Africa. But if Adria was part of Africa, then the paleolatitude anomaly of Adria relative to Europe translated into a huge crustal misfit of Gondwana relative to Laurasia when these landmasses were forced into a classic Wegenerian Pangea as typified by the Bullard fit of the circum-Atlantic continents. This crustal misfit between Gondwana and Laurasia was shown to persist in the ever-growing paleomagnetic database even when data from Adria were provisionally excluded as non-cratonic in nature. Various solutions were offered that ultimately involved placing Gondwana to the east (allowing it to be more northerly) relative to Laurasia and envisaging a dextral shear occurring in the Tethys (Mediterranean) realm between these supercontinental landmasses. This shear or transformation was initially thought to occur as a continuum over the course of the Mesozoic–Cenozoic (the so-called ‘Tethys Twist’) but soon afterwards when plate tectonics came into play and limited the younger extent, as a discrete event during the post-Triassic, Triassic or most probably – as in the latest and preferred reconstructions – the Permian between a configuration of Pangea termed B – with the northwestern margin of Africa against southern Europe – to a configuration termed Pangea A-2, with the northwestern margin of Africa against eastern North America, that is more proximal in shape to the classic Pangea A-1 that started fragmenting in the Jurassic with the opening of the Atlantic Ocean. The Permian timing and presumed locus of the ~2300 km dextral shear is supported by rotated tectonic domains in Sardinia and elsewhere along the interface between Lauarasia and Gondwana. The concept of Pangea B and its transformation into Pangea A developed therefore in close conjunction with the concept and paleomagnetic support of Adria as a promontory of Africa, and has ramifications to many aspects of tectonics, climate change and biogeography yet to be explored.","PeriodicalId":54451,"journal":{"name":"Rivista Italiana Di Paleontologia E Stratigrafia","volume":"125 1","pages":"249-269"},"PeriodicalIF":2.3,"publicationDate":"2019-03-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"47819777","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"NORIAN AMMONOIDS FROM THE NAYBAND FORMATION (IRAN) AND THEIR BEARING ON LATE TRIASSIC SEDIMENTARY AND GEODYNAMIC HISTORY OF THE IRAN PLATE","authors":"L. Krystyn, M. Balini, B. Aghababalou, V. Hairapetian","doi":"10.13130/2039-4942/11412","DOIUrl":"https://doi.org/10.13130/2039-4942/11412","url":null,"abstract":"A Middle Norian (Alaunian 2) ammonoid assemblage from north of Esfahan (Central Iran) is described and chronostratigraphically evaluated. Formerly known as Distichites fauna, it represents a geographically widely distributed and stratigraphically important fossil level in the lower part of the Upper Triassic Nayband Formation. The new distichitid ammonoid genus Mesodistichites with the new species M. evolutus are introduced; additional faunal members are Noridiscites nodosus n. sp. and the leiostracean Stenarcestes diogenis and Pinacoceras cf. imperator. The Nayband Formation of the Esfahan region, belonging to the Zefreh-Soh Facies, is lithostratigraphically emended to contain three formally introduced members (Parsefid, Venher and Niazmargh members), which are all of Norian age, whereas Rhaetian sediments are missing. These members are correlated with Norian lithostratigraphic units of the type sequence of the formation in Nayband, eastern Iran. Integration of all Iranian Nayband data allows the recognition of three 3rd order sequences within this formation and the proof of a major pre- or syn-Jurassic unconformity across Central Iran and the Central-East Iranian Microcontinent (CEIM) related to the Main-Cimmerian event. Because of the latter and of major lithostratigraphic and facial differences, we propose to exclude the Nayband Formation from the Shemshak Group. A careful review of the available biostratigraphic data from northern and southern Iran let us further assume that the collision of the Iran Plate with Eurasia occurred in the later Early Carnian and was concurrent to the onset of the Carnian Humid Episode.","PeriodicalId":54451,"journal":{"name":"Rivista Italiana Di Paleontologia E Stratigrafia","volume":"125 1","pages":"231-248"},"PeriodicalIF":2.3,"publicationDate":"2019-03-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"44043524","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"BAJOCIAN LISSOCERATINAE (HAPLOCERATOIDEA, AMMONITIDA) FROM THE MEDITERRANEAN-CAUCASIAN SUBREALM","authors":"G. Pavia, S. Fernández-López","doi":"10.13130/2039-4942/11058","DOIUrl":"https://doi.org/10.13130/2039-4942/11058","url":null,"abstract":"A revision of the Bajocian Lissoceratinae is presented. The study of a huge quantity of lissoceratins from different sites of the Western Tethys (Northwest European, Sub-Mediterranean and Mediterranean bioprovinces) within the Tethys-Panthalassa Realm provided data useful to implement the systematics of these almost neglected, never deeply analysed ammonites. Two genera, Lissoceras Bayle and Semilissoceras n. gen., are described with 16 species, among which 8 (four dimorphic, three macroconchiate and one microconchiate) new species: L. submediterraneum, L. maizetense, L. ovale, L. sturanii, L. maerteni, S. ellipticum, S. turgidulum and S. costellatum. The neotype of L. oolithicum is established. The microconchiate genus Microlissoceras is regarded as the junior synonym of the macroconchiate Lissoceras. Taxa are discussed according to four groups that gather taxa sharing common morpho-structural features. They roughly relate to successive biochronostratigraphical intervals within the Bajocian Stage and are headed by species largely known in literature: S. semicostulatum (Buckman) with suboval to compressed whorl section and rectiradiate ribs on the outer half flank; L. oolithicum (d’Orbigny) with subtriangular to ovate whorl section, large and depressed ventral saddle, large suspensive lobe; L. haugi Sturani with ovate to globular whorl section, narrow lobe E and suspensive lobe; and L. psilodiscus (Schloenbach) with highly compressed whorl section, high ventral saddle and narrow lobe E. The phyletic relation of Lissoceras and Semilissoceras to the Aalenian-Bajocian Bradfordia-group is discussed, regarding Semilissoceras as the known stem-taxon on which the subfamily Lissoceratinae bases.","PeriodicalId":54451,"journal":{"name":"Rivista Italiana Di Paleontologia E Stratigrafia","volume":" ","pages":""},"PeriodicalIF":2.3,"publicationDate":"2019-03-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"48391157","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"THE LOWER BAJOCIAN GAETANI LEVEL: LITHOSTRATIGRAPHIC MARKER OF A POTENTIAL OCEANIC ANOXIC EVENT","authors":"E. Erba, G. Gambacorta, M. Tiepolo","doi":"10.13130/2039-4942/11390","DOIUrl":"https://doi.org/10.13130/2039-4942/11390","url":null,"abstract":"In this paper we document in detail the transition from the Rosso Ammonitico Lombardo to the Radiolarites outcropping at Alpe Turati (Albavilla, Como) in the Lombardy Basin of the Southern Alps. At this location, the Jurassic succession was deposited on a flank of the Corni di Canzo paleohigh: as found on other structural shallow areas, the upper Pliensbachian–Toarcian–Aalenian interval is represented by pseudonodular to nodular marly limestones of the Morbio Limestone and Rosso Ammonitico Lombardo units. In the early Bajocian deposition of biosiliceous sediments is ubiquitously testified by the Radiolarites (Bajocian–Callovian). At Alpe Turati, the interval immediately below the basal green member of the latter unit, consists of a 10 cm thick black shale, pointing to oxygen-depleted bottom waters. Dysoxia persisted during the Bajocian–Callovian as evidenced by dark green stratified cherts. Calcareous nannofossil biostratigraphy indicate an early Bajocian age for the black shale that correlates with the core of a distinct C isotopic negative anomaly also known in various basins outside the Southern Alps. Very dark grey to black lithologies of early Bajocian age have been documented in a few sections from Poland, Corsica and Morocco. We name Gaetani Level the black shale interval in recognition of Maurizio Gaetani’s pioneering and extensive work on Jurassic sedimentary successions of the Lombardy Basin. The association of the Gaetani black shale with a C isotopic anomaly suggests that it could be the sedimentary record of an Oceanic Anoxic Event whose regional to global extension must be ascertained.","PeriodicalId":54451,"journal":{"name":"Rivista Italiana Di Paleontologia E Stratigrafia","volume":"125 1","pages":"219-230"},"PeriodicalIF":2.3,"publicationDate":"2019-03-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"44080142","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"LANTERNFISH OTOLITHS (TELEOSTEI, MYCTOPHIDAE) FROM THE PLIOCENE AND PLEISTOCENE OF JAPAN","authors":"W. Schwarzhans, Fumio Ohe","doi":"10.13130/2039-4942/11670","DOIUrl":"https://doi.org/10.13130/2039-4942/11670","url":null,"abstract":"Myctophid otoliths are the most common fossil otoliths to be found in Neogene deep water sediments below approximately 200 m depositional depth. The southeastern part of Japan is particularly rich in such locations. Here we describe myctophid otoliths from late Pliocene to early Pleistocene locations on Okinawa, Shikoku and central Honshu and review previous publications. These faunas represent the tropical to subtropical faunal zone, which was established as the “Kakegawa Fauna” based on molluscs. A total of 37 species are recognized by means of otoliths in the family Myctophidae and one in the family Neoscopelidae. Two species are recorded in open nomenclature, 23 as persistent extant species and 12 extinct species, thereof 7 new. The new species are: Diaphus caurus, Diaphus endoi, Diaphus kakegawaensis, Diaphus nafpaktitisi, Diaphus noboriensis, Symbolophorus moriguchii and Notoscopelus praejaponicus. Two very large myctophid otoliths (>8 mm length) from the putative Piacenzian of Fiji are also described, one being a new species - Diaphus grebneffi - representing the largest by far otolith associated with the So-group of species in the genus so far known. The myctophid otolith assemblage is characterized by an abundance of large specimens (>5 mm length) which certainly derived from fully adult fishes and represents the biggest association of its kind described so far from the fossil record of the northern Pacific. Its composition is interpreted for stratigraphic and environmental purposes. The occurrence of a few, rare species with links to coeval finds in the Caribbean exemplify the potential of myctophid otoliths for supraregional biostratigraphic purposes. The comparison of the fossil assemblage with the distribution in the Recent supports the concept of a warm “Kakegawa Fauna” as established by molluscs and at the same time documents that certain tropical species reached further north during the late Pliocene than they do today.","PeriodicalId":54451,"journal":{"name":"Rivista Italiana Di Paleontologia E Stratigrafia","volume":"125 1","pages":""},"PeriodicalIF":2.3,"publicationDate":"2019-01-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"66212701","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"PALEOGEOGRAPHIC NORTHEASTERN LIMITS OF APHRODINA DUTRUGEI (COCQUAND, 1862) (HETERODONTA, BIVALVIA) FROM THE CENOMANIAN OF THE ARABIAN PLATFORM","authors":"I. Hoşgör, İ. Yılmaz","doi":"10.13130/2039-4942/11773","DOIUrl":"https://doi.org/10.13130/2039-4942/11773","url":null,"abstract":"The finding of Aphrodina dutrugei (Cocquand, 1862) in a rich collection of non-rudist bivalve fauna from the famous Cenomanian-Turonian carbonates of the Afro-Arabian Plate has permitted the reevaluation of venerid bivalves during an important period of their evolution. The genus Aphrodina Conrad, 1869, Family Veneridae Rafinesque, 1815, embraces several Tethyan Cretaceous species that range from Cenomanian to Santonian. During the late Albian-Maastrichtian ‘Aphrodiniid’ venerids, were distributed along the western margin of the Atlantic (North and South America), the Afro-Arabian Plate (Jordan, SE Turkey, Morocco, Algeria and Egypt), the eastern Tethys, and the Southern Ocean (India, Japan, western Australia and New Zealand). They are also known in the Turonian-Santonian Trans-Saharan Seaway (Gabon). Until now these fossils have been unknown in any Upper Cretaceous localities of southeastern Turkey. In this paper, we report the first record of one of the most common and widespread shallow infaunal species, Aphrodina dutrugei, in the Cenomanian Derdere Formation in the Mardin-Mazidagi area, SE Turkey, which is in the extreme northeastern part of its known range.","PeriodicalId":54451,"journal":{"name":"Rivista Italiana Di Paleontologia E Stratigrafia","volume":"125 1","pages":""},"PeriodicalIF":2.3,"publicationDate":"2019-01-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"66212746","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"GLOBIGAETANIA ANGULATA GEN. N. SP. N. (GLOBIVALVULININAE, FORAMINIFERA) FROM THE WORDIAN (MIDDLE PERMIAN) OF NW IRAN","authors":"Valerio Gennari, R. Rettori","doi":"10.13130/2039-4942/11054","DOIUrl":"https://doi.org/10.13130/2039-4942/11054","url":null,"abstract":"A new small biseriamminoid foraminifer, Globigaetania angulata gen. n. sp. n., is here described from the Wordian of a Permian –Triassic sedimentary succession of NW Iran (Zal and Poldasht stratigraphic sections). The new taxon, dedicated to Prof. Maurizio Gaetani, is characterised by peculiar morphology, coiling, and structures that are characteristic of the subfamily Globivalvulininae Reitlinger, 1950, family Globivalvulinidae Reitlinger, 1950, superfamily Biseriamminoidea Chernysheva, 1941. The introduction of the new taxon contributes to the knowledge of the systematics and evolution of the Palaeozoic biserial microgranular foraminifera.","PeriodicalId":54451,"journal":{"name":"Rivista Italiana Di Paleontologia E Stratigrafia","volume":"125 1","pages":""},"PeriodicalIF":2.3,"publicationDate":"2019-01-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"66212463","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"LOWER TRIASSIC (INDUAN TO OLENEKIAN) CONODONTS, FORAMINIFERA AND BIVALVES FROM THE AL MAMALIH AREA, DEAD SEA, JORDAN: CONSTRAINTS ON THE P-T BOUNDARY","authors":"J. Powell, A. Nicora, M. Perri, R. Rettori, R. Posenato, M. Stephenson, A. Masri, Letizia M. Borlenghi, Valerio Gennari","doi":"10.13130/2039-4942/11270","DOIUrl":"https://doi.org/10.13130/2039-4942/11270","url":null,"abstract":"Upper Permian to Lower Triassic successions exposed in the Al Mamalih area, east of the Dead Sea, Jordan record the transition between the alluvial Umm Irna Formation (Upper Permian) and the overlying shallow marine Ma’in Formation (Lower Triassic). The Permian-Triassic boundary is constrained either within a hiatus represented by a sequence boundary between these formations or within ca 15 m of shallow marine beds overlying the boundary. Above the sequence boundary reddened, shallow-marine beds (Himara Member) mark the initial Triassic (presumed early Induan) marine transgression (Himara Member). Absence of both body fossils and vertical infaunal burrows indicates low-diversity, ecosystems following the Permian-Triassic extinction event. A gradational upward increase in grey, green and yellow siltstones beds (Nimra Member), accompanied by a concomitant increase in bioturbation (surface traces and infaunal vertical burrows), bivalves and brachiopods, stromatolites, conchostracans and lingulids in the lower part of the Nimra Member indicates colonisation of the substrate under shallow marine conditions during the recovery phase. Shallow-water carbonates in the Nimra Member yielded an abundant, low diversity assemblage of conodonts (e.g. Hd. aequabilis and H. agordina) and a foraminifera assemblage (Postcladella gr. kalhori-Earlandia spp.-Ammodiscus jordanensis n. sp.) that are interpreted as euryhaline recovery taxa that characterise the the mid-late Induan. Abundant new material has allowed revision of the conodont apparatus and the foraminifera include a new species Ammodiscus jordanensis n. sp. of Induan age. The discovery of the bivalves Claraia bittneri (C. aurita group) and Eumorphotis multiformis is worthy to note. Upper Permian alluvial lithofacies (Jordan) pass basinwards, about 50 km to the northwest, to coeval shallow marine siliciclastic and carbonates in the Negev and Mediterranean coast of Israel.","PeriodicalId":54451,"journal":{"name":"Rivista Italiana Di Paleontologia E Stratigrafia","volume":"125 1","pages":""},"PeriodicalIF":2.3,"publicationDate":"2019-01-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"66212559","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

{"title":"PALEOECOLOGY AND FUNCTIONAL MORPHOLOGY OF THE PERMIAN LYTTONIID BRACHIOPOD PIRGULIA","authors":"Daniel J. Stadtmauer, Susan H. Butts","doi":"10.13130/2039-4942/12172","DOIUrl":"https://doi.org/10.13130/2039-4942/12172","url":null,"abstract":"The lyttoniid brachiopods of the Permian exhibit a unique valve morphology: a branched lobate structure takes the form of the dorsal valve. In one group of lyttoniids, the genus Pirgulia, the ventral valve wraps around to form a cone that fully encloses the lobate structure. This has consequences for the dynamics of water flow and mode of life possible for these heteromorphic brachiopods. Here, we describe the skeletal microstructure and morphology of Pirgulia collected from the Upper Permian Sosio Limestone megablocks of Sicily and housed at the Yale Peabody Museum. We reconstruct the paleoecology of Pirgulia, characterizing it as semi-infaunal in soft sediment. By analogy to Richthofenia, the conical ventral valve and flapping dorsal valve functional morphology could have resisted fouling and assisted feeding in this environment. By comparison with the functional morphology of Pirgulia with other lyttoniids and richthofenids, we propose a revised mode of life for this genus, which involves adaptation to secondary soft-bottom substrates and support by sediment sticking. Despite constraints to the fundamental brachiopod body plan, modification of the valves in Pirgulia to achieve a conical morphology allowed it to inhabit a paleoecological niche distinct from that of other reef-building lyttoniids.","PeriodicalId":54451,"journal":{"name":"Rivista Italiana Di Paleontologia E Stratigrafia","volume":"125 1","pages":""},"PeriodicalIF":2.3,"publicationDate":"2019-01-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"66213041","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}