{"title":"探索捕食者-猎物动力学:整合竞争对手捕食者,利用改进的Beddington-DeAngelis功能反应收集猎物的延迟和恐惧效应","authors":"Anuj Kumar Umrao, Prashant K. Srivastava","doi":"10.1016/j.nonrwa.2025.104391","DOIUrl":null,"url":null,"abstract":"<div><div>Selective harvesting is a vital strategy for controlling over-exploitation and protecting the incidental killing of juvenile species. To ensure that the individuals reach a suitable age or size before being harvested, a specific time delay, known as harvesting-induced delay, should be maintained. Also, predators can indirectly slow down the growth rate of prey by inducing fear in them, which affects their behaviour and reproduction. In this work, we study a predator–prey model to examine the impact of predator selective (delayed) harvesting in the presence of fear in prey species and the interspecific competition of predator with the competitive predator. It is assumed that the density of the competitive predator is constant and both predators induce fear in prey. We determine the conditions related to the existence of transcritical and Hopf bifurcations for the non-delay case, and various delay-induced scenarios via Hopf bifurcation. We also numerically explore the impact of delayed harvesting on the system dynamics by simultaneously varying the harvesting effort, fear level, and the interspecific competition with competitive predator in bi-parametric planes. It is observed that the harvesting delay can result in stability invariance, instability invariance, stability change, instability switching, and stability switching phenomena under varied parametric conditions. Further, when the harvesting delay is fixed, the equilibrium point experiences instability invariance and instability change, instability change and instability switching, and instability invariance, stability change and stability switching with respect to the harvesting effort, the interspecific competition, and the fear level, respectively. Our findings indicate that regulated selective harvesting of predator, and a moderate level of fear in prey and interspecific competition with the competitive predator are essential for maintaining stability and coexistence in the ecosystem. The rich and complex dynamics holds significance from a biological viewpoint and may potentially impact the population management strategies.</div></div>","PeriodicalId":49745,"journal":{"name":"Nonlinear Analysis-Real World Applications","volume":"86 ","pages":"Article 104391"},"PeriodicalIF":1.8000,"publicationDate":"2025-04-28","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":"0","resultStr":"{\"title\":\"Exploring predator–prey dynamics: Integrating competitor predators, harvesting delay and fear effect on prey with a modified Beddington–DeAngelis functional response\",\"authors\":\"Anuj Kumar Umrao, Prashant K. Srivastava\",\"doi\":\"10.1016/j.nonrwa.2025.104391\",\"DOIUrl\":null,\"url\":null,\"abstract\":\"<div><div>Selective harvesting is a vital strategy for controlling over-exploitation and protecting the incidental killing of juvenile species. To ensure that the individuals reach a suitable age or size before being harvested, a specific time delay, known as harvesting-induced delay, should be maintained. Also, predators can indirectly slow down the growth rate of prey by inducing fear in them, which affects their behaviour and reproduction. In this work, we study a predator–prey model to examine the impact of predator selective (delayed) harvesting in the presence of fear in prey species and the interspecific competition of predator with the competitive predator. It is assumed that the density of the competitive predator is constant and both predators induce fear in prey. We determine the conditions related to the existence of transcritical and Hopf bifurcations for the non-delay case, and various delay-induced scenarios via Hopf bifurcation. We also numerically explore the impact of delayed harvesting on the system dynamics by simultaneously varying the harvesting effort, fear level, and the interspecific competition with competitive predator in bi-parametric planes. It is observed that the harvesting delay can result in stability invariance, instability invariance, stability change, instability switching, and stability switching phenomena under varied parametric conditions. Further, when the harvesting delay is fixed, the equilibrium point experiences instability invariance and instability change, instability change and instability switching, and instability invariance, stability change and stability switching with respect to the harvesting effort, the interspecific competition, and the fear level, respectively. Our findings indicate that regulated selective harvesting of predator, and a moderate level of fear in prey and interspecific competition with the competitive predator are essential for maintaining stability and coexistence in the ecosystem. The rich and complex dynamics holds significance from a biological viewpoint and may potentially impact the population management strategies.</div></div>\",\"PeriodicalId\":49745,\"journal\":{\"name\":\"Nonlinear Analysis-Real World Applications\",\"volume\":\"86 \",\"pages\":\"Article 104391\"},\"PeriodicalIF\":1.8000,\"publicationDate\":\"2025-04-28\",\"publicationTypes\":\"Journal Article\",\"fieldsOfStudy\":null,\"isOpenAccess\":false,\"openAccessPdf\":\"\",\"citationCount\":\"0\",\"resultStr\":null,\"platform\":\"Semanticscholar\",\"paperid\":null,\"PeriodicalName\":\"Nonlinear Analysis-Real World Applications\",\"FirstCategoryId\":\"100\",\"ListUrlMain\":\"https://www.sciencedirect.com/science/article/pii/S146812182500077X\",\"RegionNum\":3,\"RegionCategory\":\"数学\",\"ArticlePicture\":[],\"TitleCN\":null,\"AbstractTextCN\":null,\"PMCID\":null,\"EPubDate\":\"\",\"PubModel\":\"\",\"JCR\":\"Q1\",\"JCRName\":\"MATHEMATICS, APPLIED\",\"Score\":null,\"Total\":0}","platform":"Semanticscholar","paperid":null,"PeriodicalName":"Nonlinear Analysis-Real World Applications","FirstCategoryId":"100","ListUrlMain":"https://www.sciencedirect.com/science/article/pii/S146812182500077X","RegionNum":3,"RegionCategory":"数学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":null,"EPubDate":"","PubModel":"","JCR":"Q1","JCRName":"MATHEMATICS, APPLIED","Score":null,"Total":0}
Exploring predator–prey dynamics: Integrating competitor predators, harvesting delay and fear effect on prey with a modified Beddington–DeAngelis functional response
Selective harvesting is a vital strategy for controlling over-exploitation and protecting the incidental killing of juvenile species. To ensure that the individuals reach a suitable age or size before being harvested, a specific time delay, known as harvesting-induced delay, should be maintained. Also, predators can indirectly slow down the growth rate of prey by inducing fear in them, which affects their behaviour and reproduction. In this work, we study a predator–prey model to examine the impact of predator selective (delayed) harvesting in the presence of fear in prey species and the interspecific competition of predator with the competitive predator. It is assumed that the density of the competitive predator is constant and both predators induce fear in prey. We determine the conditions related to the existence of transcritical and Hopf bifurcations for the non-delay case, and various delay-induced scenarios via Hopf bifurcation. We also numerically explore the impact of delayed harvesting on the system dynamics by simultaneously varying the harvesting effort, fear level, and the interspecific competition with competitive predator in bi-parametric planes. It is observed that the harvesting delay can result in stability invariance, instability invariance, stability change, instability switching, and stability switching phenomena under varied parametric conditions. Further, when the harvesting delay is fixed, the equilibrium point experiences instability invariance and instability change, instability change and instability switching, and instability invariance, stability change and stability switching with respect to the harvesting effort, the interspecific competition, and the fear level, respectively. Our findings indicate that regulated selective harvesting of predator, and a moderate level of fear in prey and interspecific competition with the competitive predator are essential for maintaining stability and coexistence in the ecosystem. The rich and complex dynamics holds significance from a biological viewpoint and may potentially impact the population management strategies.
期刊介绍:
Nonlinear Analysis: Real World Applications welcomes all research articles of the highest quality with special emphasis on applying techniques of nonlinear analysis to model and to treat nonlinear phenomena with which nature confronts us. Coverage of applications includes any branch of science and technology such as solid and fluid mechanics, material science, mathematical biology and chemistry, control theory, and inverse problems.
The aim of Nonlinear Analysis: Real World Applications is to publish articles which are predominantly devoted to employing methods and techniques from analysis, including partial differential equations, functional analysis, dynamical systems and evolution equations, calculus of variations, and bifurcations theory.