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Citrus have been cultivated more than 4,000 years and have become globally valued as fresh fruit and for making juice. Citrus fruits contain many types of antioxidants, including a diverse array of carotenoids, which contribute to their colors. The cover shows cross-sections of citrus fruits arranged in a spiral pattern and displaying a stunning array of colors, from white and pale yellow to grapefruit pink, blood red, golden yellow, and vibrant orange. These slices, varying in size and hue highlight the richness and diversity of citrus fruits and emphasizing the vibrant palette of citrus carotenoids. Sun et al. (pages 294-310) identified CitZAT4 as a key regulator of carotenoid accumulation, providing valuable strategies for breeding citrus varieties with enhanced quality and nutritional value, thereby boosting the economic benefits of the citrus industry.