Supplementation of Yearling Steers Grazing Fertilized and Unfertilized Northern Plains Rangeland

J. Karn, R. Lorenz
{"title":"Supplementation of Yearling Steers Grazing Fertilized and Unfertilized Northern Plains Rangeland","authors":"J. Karn, R. Lorenz","doi":"10.2307/3897979","DOIUrl":null,"url":null,"abstract":"Supplementation studies were conducted with yearling steers on a silty range site in central North Dakota, where yearly precipitation averaged 380 to 410 mm. The studies were conducted for 3 summers on both fertilized (45 kg N/ha) and unfertilized native pastures. Animal performance was compared to seasonal changes in the chemical composition of pasture samples collected with esophageal-fistulated steers. Chemical composition differences between diet samples from the fertilized and unfertilized pastures were inconsistent, but generally protein was higher and acid detergent fiber lower on the fertilized pasture. Supplementation with barley in the early summer resulted in little benefit, but supplementation with barley in the late summer, especially when pasture digestibility (in vitro) dropped to 50 to 52%, was beneficial on both the fertilized and unfertilized pastures. However, the response was not consistent between years. Barley supplementation appeared to be economically viable, but the feasibility of this practice will vary from year to year, depending on the price of barley relative to the price of steers. The results of protein supplementation were more erratic, possibly because of differences in precipitation patterns and hence plant growth between years. Native rangelands in the Northern Great Plains support good summer weight gains on yearling steers. Therefore, little consideration has been given to use of supplements which might produce still greater and more efficient gains. Rogler and Lorenz (1965) have shown that beef cattle carrying capacity of the Northern Great Plains mixed grass prairie can be approximately doubled with appropriate nitrogen fertilization. However, fertilization was shown to favor the growth of cool-season midgrasses, particularly western wheatgrass (Agropyron smithii), with an associated reduction in the stand of blue grama (Boutelouagracilis), a warm-season shortgrass (Lorenz and Rogler 1972). This shift in species composition alters the seasonal growth pattern and may affect the nutritional value of the grazing animal’s diet. There is limited information available on the chemical composition of native forages in the Northern Great Plains, especially with respect to changes during the grazing season. Information available on the value of supplementing yearling steers grazing native forage is also limited. However, Raleigh (1970) reported positive results from a supplementation scheme developed for use in eastern Oregon. Both crude protein (CP) and energy were supplemented as required to complement the diet of yearling steers grazing crested wheatgrass (Agropyron desertorum). A beneficial effect from energy supplementation of steers grazing spring native range in eastern Colorado has been reported by Denham (1977). These results suggested that CP and/or energy supplementation might also be beneficial to grazing steers in the Northern Plains. Thus the objectives of this study were: (I) to determine the chemiAuthors are research animal scientist and research agronomist, USDA-AR& Northern Great Plains Research Laboratory, P.O. Box 459, Mandan, N.D. 58554. Authors wish toacknowledge Mr. Charles Graham for hisassistance with statistical procedures and Mr. Richard Huppler and Mrs. Margo Dockter for their techncial assistance. Manuscript received May I I, 1980. cal composition of esophageal fistula diet collections taken from fertilized and unfertilized native range throughout the grazing season, and (2) to study the effect of CP and energy supplementation on the performance of yearling steers grazing these rangelands. Materials and Methods Summer supplementation studies were undertaken in 1977, 1978, and 1979 on a Northern Great Plains silty range site in the 380-410 mm rainfall area of central North Dakota. Predominate forage species were western wheatgrass (Agropyron smithii). prairie junegrass (Koeleria cristata), blue gmma (Boutelouagracilis), needleandthread (Stipa comata), green needlegrass (Stipa virid&), and upland sedges (Carex spp.). Big bluestem (Andropogon gerardii) was present in low areas and along intermittent waterways. The experimental site had been previously separated into 2 pastures. The smaller of these (14.2 hectares) had been fertilized annually for 20 years with 45 kg nitrogen per hectare (45-N). This practice was continued during these studies. The other pasture contained 28.4 hectares and had not been fertilized (O-N). The same number of steers were grazed on each pasture based on results of work by Rogler and Lorenz (1965), which demonstrated that a hectare of fertilized rangeland would support twice as many steers as a hectare of unfertilized rangeland. Five esophageal-fistulated steers were used to collect weekly diet samples from each pasture in 1977 and 1978. The 1977 study was conducted from mid-May until mid-September and the 1978 study from mid-May until mid-October. In 1979 diet samples were collected every second week from mid-May to mid-October using 6 esophageal-fistulated steers. Diet samples were collected from both pastures the same day by using the same steers. To avoid a possible pasture bias, samples were collected one week from the 45-N pasture first and the next week from the O-N pasture first, Obioha et al. (1970), reported slight but significant differences in nitrogen level between morning and evening forage samples collected via esophageal fistula. The difference was attributed to differences in grazing selectivity due to hunger. In our study samples were collected from pastures in the morning, with samples collected from the second pasture immediately following the first. Esophageal-fistulated steers were maintained on an adjacent O-N native pasture and were not supplemented. They were kept off feed the night before sampling to facilitate diet collections. Diet samples collected by individual steers were dried to a constant weight at SO“ C in shallow pans in a forced draft oven then ground throught a l-mm screen in a Wiley mill. Chemical analysis included Kjeldahl nitrogen, in vitro digestible organic matter (Tilley and Terry 1963, Moore and Mott 1974), neutral detergent fiber (Van Soest and Wine 1967), acid detergent fiber (ADF) and lignin (Van Soest and Wine 1968). In 1978 and 1979 diet samples from individual steers were processed the same as in 1977. Kjeldahl nitrogen was measured on each sample, but the other chemical analyses were performed on a composite sample for each pasture JOURNAL OF RANGE MANAGEMENT 36(l), January 1963 41 Table 1. Chemical comypition of forage samples collected via esopbagel fiitulated steers during tbe summer grazing season from native range receiving 45-N and O-N (1977). Month IVDOM % Pasture 45-N O-N CP NDF ADF % of D.M. Lignin % of D.M. % of D.M. % of D.M. Pasture Pasture Pasture Pasture Mean 45-N O-N Mean 45-N O-N Mean 45-N O-N Mean 45-N O-N Mean May’ 66.2 62.4 64.3 19.1 12.8 16.0 56.6 66.6 61.6 33.5 46.8 37.2 7.6 6.5 7.0 June 59.7 57.4 58.6 13.4 10.5 12.0 69.4 JO.6 70.0 39.1 44.3 41.7 6.4 8.2 7.3 July 56.6 53.6 55.1 12.7 8.8 10.8 10.6 JO.6 70.6 41.5 44.6 43.0 7.2 7.7 7.4 August 58.2 52.5 55.4 12.2 7.9 10.0 67.3 66.4 66.8 42.2 46.6 44.4 7.4 9.1 8.2 September 60.3 55.9 58.1 16.2 12.6 14.4 64.5 66. I 65.3 39.1 46.8 43.0 6.8 9.6 8.2 mean 60.2” 56.4’ 58.3 14.7” 10.5b 12.6 65.7” 68.lb 66.9 39. I” 44.6b 41.9 7.1’ 8.2b 7.6 ‘Pasture means with the same heading and different superscripts differ significantly (K.05). *Each value isan average of20samples(4dates by5steers)except September, whichcontains only3samplingdatesand May whichcontains 19samplesfor the45-N pastureand I8 for the O-N pasture. ‘Contains June 2 sampling date. for each sampling date. Data wereanalyzed by collection period by using either an arithmetic mean of data from individual diet samples or data from the chemical analysis of a composite sample. equipment to facilitate supplementation when the study was initiated. 1977 Study Twenty yearling Hereford steers weighing approximately265 kg were randomly assigned to either the 45-N or O-N pasture. Five steers on each pasture were then randomly assigned to one of the following treatments: (I) no supplement, or (2) .9 kg dry rolled barley containing 2.7 Meal DE and 119 gm CP. Steers were corralled each morning before 1000 hours CDT and those that received supplement were individually fed. Refused supplement was weighed and subtracted from the amount offered. The experiment was started June 1 and terminated July 22 because of a severe drought and consequent lack of forage, which forced removal of the steers from the pasture. All steers had continual access to a 50-50 mixture of salt and dicalcium phosphate. Weight gains for each pasture, each period, and each year were analyzed separately, then the two years of ES data and the two years of LS data were analyzed as randomized complete blocks. Treatment differences were determined by Duncan’s multiple range test and were considered statistically significant at the 5% level of probability.","PeriodicalId":16918,"journal":{"name":"Journal of Range Management","volume":"101 1","pages":"41"},"PeriodicalIF":0.0000,"publicationDate":"2006-06-19","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":"3","resultStr":null,"platform":"Semanticscholar","paperid":null,"PeriodicalName":"Journal of Range Management","FirstCategoryId":"1085","ListUrlMain":"https://doi.org/10.2307/3897979","RegionNum":0,"RegionCategory":null,"ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":null,"EPubDate":"","PubModel":"","JCR":"","JCRName":"","Score":null,"Total":0}
引用次数: 3

Abstract

Supplementation studies were conducted with yearling steers on a silty range site in central North Dakota, where yearly precipitation averaged 380 to 410 mm. The studies were conducted for 3 summers on both fertilized (45 kg N/ha) and unfertilized native pastures. Animal performance was compared to seasonal changes in the chemical composition of pasture samples collected with esophageal-fistulated steers. Chemical composition differences between diet samples from the fertilized and unfertilized pastures were inconsistent, but generally protein was higher and acid detergent fiber lower on the fertilized pasture. Supplementation with barley in the early summer resulted in little benefit, but supplementation with barley in the late summer, especially when pasture digestibility (in vitro) dropped to 50 to 52%, was beneficial on both the fertilized and unfertilized pastures. However, the response was not consistent between years. Barley supplementation appeared to be economically viable, but the feasibility of this practice will vary from year to year, depending on the price of barley relative to the price of steers. The results of protein supplementation were more erratic, possibly because of differences in precipitation patterns and hence plant growth between years. Native rangelands in the Northern Great Plains support good summer weight gains on yearling steers. Therefore, little consideration has been given to use of supplements which might produce still greater and more efficient gains. Rogler and Lorenz (1965) have shown that beef cattle carrying capacity of the Northern Great Plains mixed grass prairie can be approximately doubled with appropriate nitrogen fertilization. However, fertilization was shown to favor the growth of cool-season midgrasses, particularly western wheatgrass (Agropyron smithii), with an associated reduction in the stand of blue grama (Boutelouagracilis), a warm-season shortgrass (Lorenz and Rogler 1972). This shift in species composition alters the seasonal growth pattern and may affect the nutritional value of the grazing animal’s diet. There is limited information available on the chemical composition of native forages in the Northern Great Plains, especially with respect to changes during the grazing season. Information available on the value of supplementing yearling steers grazing native forage is also limited. However, Raleigh (1970) reported positive results from a supplementation scheme developed for use in eastern Oregon. Both crude protein (CP) and energy were supplemented as required to complement the diet of yearling steers grazing crested wheatgrass (Agropyron desertorum). A beneficial effect from energy supplementation of steers grazing spring native range in eastern Colorado has been reported by Denham (1977). These results suggested that CP and/or energy supplementation might also be beneficial to grazing steers in the Northern Plains. Thus the objectives of this study were: (I) to determine the chemiAuthors are research animal scientist and research agronomist, USDA-AR& Northern Great Plains Research Laboratory, P.O. Box 459, Mandan, N.D. 58554. Authors wish toacknowledge Mr. Charles Graham for hisassistance with statistical procedures and Mr. Richard Huppler and Mrs. Margo Dockter for their techncial assistance. Manuscript received May I I, 1980. cal composition of esophageal fistula diet collections taken from fertilized and unfertilized native range throughout the grazing season, and (2) to study the effect of CP and energy supplementation on the performance of yearling steers grazing these rangelands. Materials and Methods Summer supplementation studies were undertaken in 1977, 1978, and 1979 on a Northern Great Plains silty range site in the 380-410 mm rainfall area of central North Dakota. Predominate forage species were western wheatgrass (Agropyron smithii). prairie junegrass (Koeleria cristata), blue gmma (Boutelouagracilis), needleandthread (Stipa comata), green needlegrass (Stipa virid&), and upland sedges (Carex spp.). Big bluestem (Andropogon gerardii) was present in low areas and along intermittent waterways. The experimental site had been previously separated into 2 pastures. The smaller of these (14.2 hectares) had been fertilized annually for 20 years with 45 kg nitrogen per hectare (45-N). This practice was continued during these studies. The other pasture contained 28.4 hectares and had not been fertilized (O-N). The same number of steers were grazed on each pasture based on results of work by Rogler and Lorenz (1965), which demonstrated that a hectare of fertilized rangeland would support twice as many steers as a hectare of unfertilized rangeland. Five esophageal-fistulated steers were used to collect weekly diet samples from each pasture in 1977 and 1978. The 1977 study was conducted from mid-May until mid-September and the 1978 study from mid-May until mid-October. In 1979 diet samples were collected every second week from mid-May to mid-October using 6 esophageal-fistulated steers. Diet samples were collected from both pastures the same day by using the same steers. To avoid a possible pasture bias, samples were collected one week from the 45-N pasture first and the next week from the O-N pasture first, Obioha et al. (1970), reported slight but significant differences in nitrogen level between morning and evening forage samples collected via esophageal fistula. The difference was attributed to differences in grazing selectivity due to hunger. In our study samples were collected from pastures in the morning, with samples collected from the second pasture immediately following the first. Esophageal-fistulated steers were maintained on an adjacent O-N native pasture and were not supplemented. They were kept off feed the night before sampling to facilitate diet collections. Diet samples collected by individual steers were dried to a constant weight at SO“ C in shallow pans in a forced draft oven then ground throught a l-mm screen in a Wiley mill. Chemical analysis included Kjeldahl nitrogen, in vitro digestible organic matter (Tilley and Terry 1963, Moore and Mott 1974), neutral detergent fiber (Van Soest and Wine 1967), acid detergent fiber (ADF) and lignin (Van Soest and Wine 1968). In 1978 and 1979 diet samples from individual steers were processed the same as in 1977. Kjeldahl nitrogen was measured on each sample, but the other chemical analyses were performed on a composite sample for each pasture JOURNAL OF RANGE MANAGEMENT 36(l), January 1963 41 Table 1. Chemical comypition of forage samples collected via esopbagel fiitulated steers during tbe summer grazing season from native range receiving 45-N and O-N (1977). Month IVDOM % Pasture 45-N O-N CP NDF ADF % of D.M. Lignin % of D.M. % of D.M. % of D.M. Pasture Pasture Pasture Pasture Mean 45-N O-N Mean 45-N O-N Mean 45-N O-N Mean 45-N O-N Mean May’ 66.2 62.4 64.3 19.1 12.8 16.0 56.6 66.6 61.6 33.5 46.8 37.2 7.6 6.5 7.0 June 59.7 57.4 58.6 13.4 10.5 12.0 69.4 JO.6 70.0 39.1 44.3 41.7 6.4 8.2 7.3 July 56.6 53.6 55.1 12.7 8.8 10.8 10.6 JO.6 70.6 41.5 44.6 43.0 7.2 7.7 7.4 August 58.2 52.5 55.4 12.2 7.9 10.0 67.3 66.4 66.8 42.2 46.6 44.4 7.4 9.1 8.2 September 60.3 55.9 58.1 16.2 12.6 14.4 64.5 66. I 65.3 39.1 46.8 43.0 6.8 9.6 8.2 mean 60.2” 56.4’ 58.3 14.7” 10.5b 12.6 65.7” 68.lb 66.9 39. I” 44.6b 41.9 7.1’ 8.2b 7.6 ‘Pasture means with the same heading and different superscripts differ significantly (K.05). *Each value isan average of20samples(4dates by5steers)except September, whichcontains only3samplingdatesand May whichcontains 19samplesfor the45-N pastureand I8 for the O-N pasture. ‘Contains June 2 sampling date. for each sampling date. Data wereanalyzed by collection period by using either an arithmetic mean of data from individual diet samples or data from the chemical analysis of a composite sample. equipment to facilitate supplementation when the study was initiated. 1977 Study Twenty yearling Hereford steers weighing approximately265 kg were randomly assigned to either the 45-N or O-N pasture. Five steers on each pasture were then randomly assigned to one of the following treatments: (I) no supplement, or (2) .9 kg dry rolled barley containing 2.7 Meal DE and 119 gm CP. Steers were corralled each morning before 1000 hours CDT and those that received supplement were individually fed. Refused supplement was weighed and subtracted from the amount offered. The experiment was started June 1 and terminated July 22 because of a severe drought and consequent lack of forage, which forced removal of the steers from the pasture. All steers had continual access to a 50-50 mixture of salt and dicalcium phosphate. Weight gains for each pasture, each period, and each year were analyzed separately, then the two years of ES data and the two years of LS data were analyzed as randomized complete blocks. Treatment differences were determined by Duncan’s multiple range test and were considered statistically significant at the 5% level of probability.
北方平原牧场施肥与未施肥条件下一岁阉牛的饲粮补充
补充研究是在北达科他州中部的一个粉砂质牧场进行的,那里的年平均降水量为380至410毫米。试验在施肥(45 kg N/ hm2)和未施肥的天然牧场上进行,为期3个夏季。动物生产性能与食道瘘牛采集的牧草样品化学成分的季节变化进行了比较。施肥与未施肥牧场饲粮样品化学成分差异不一致,但施肥牧场饲粮蛋白质含量较高,酸性洗涤纤维含量较低。在初夏补饲大麦的效果不大,但在夏末补饲大麦,特别是当牧草体外消化率降至50% ~ 52%时,对施肥和未施肥的牧场都有好处。然而,不同年份的反应并不一致。大麦补充似乎在经济上是可行的,但这种做法的可行性每年都会有所不同,这取决于大麦的价格相对于阉牛的价格。补充蛋白质的结果更加不稳定,可能是因为降水模式的差异,因此不同年份的植物生长。在北部大平原的原生牧场支持良好的夏季体重增加一岁的阉牛。因此,很少考虑使用可能产生更大和更有效收益的补品。Rogler和Lorenz(1965)的研究表明,在适当的氮肥处理下,北方大平原混交草草原的肉牛承载能力可提高约一倍。然而,施肥有利于冷季中草的生长,特别是西部小麦草(Agropyron smithii),与此相关的是暖季短草蓝草(bouelouagracilis)林分的减少(Lorenz和Rogler 1972)。这种物种组成的转变改变了季节性生长模式,并可能影响食草动物饮食的营养价值。关于北方大平原原生牧草的化学成分,特别是放牧季节的变化,现有的资料有限。有关补充一岁阉牛吃本地牧草的价值的信息也很有限。然而,Raleigh(1970)报道了俄勒冈州东部使用的补充方案的积极结果。在饲粮中按需要补充粗蛋白质和能量。Denham(1977)报道了在科罗拉多州东部的春季牧场放牧的牛补充能量的有益效果。这些结果表明,CP和/或能量补充也可能对北方平原放牧的阉牛有益。因此,本研究的目的是:(1)确定化学成分,作者是研究动物科学家和研究农学家,美国农业部农业部和北方大平原研究实验室,邮编:4559号,Mandan, n.d.d 58554。作者谨感谢Charles Graham先生在统计程序上的协助,以及Richard Huppler先生和Margo Dockter夫人在技术上的协助。1980年5月1日收稿。(2)研究饲粮添加CP和能量对放牧这些牧场的阉阉阉牛生产性能的影响。材料和方法于1977年、1978年和1979年在北达科他州中部降雨量380-410毫米的北部大平原粉砂质地区进行了夏季补充研究。主要牧草种为西部小麦草(Agropyron smithii)。草原六月草(Koeleria cristata)、蓝针茅(Boutelouagracilis)、针线草(Stipa comata)、绿针茅(Stipa virid&)和高地莎草(Carex spp.)。大蓝茎(Andropogon gerardii)出现在低洼地区和间歇水道。实验场地以前被分成两个牧场。其中较小的一块(14.2公顷)每年施肥20年,每公顷45公斤氮(45- n)。在这些研究中,这种做法一直在继续。另一个牧场面积28.4公顷,未施肥(O-N)。根据Rogler和Lorenz(1965)的工作结果,在每个牧场上放牧的牛数量相同,这表明一公顷施肥的牧场能养活两倍于一公顷未施肥的牧场。1977年和1978年,5头食道瘘的阉牛每周从每个牧场采集日粮样本。1977年的研究从5月中旬进行到9月中旬,1978年的研究从5月中旬进行到10月中旬。 1979年5月中旬至10月中旬,每隔两周采集6例食管瘘患者的饮食样本。在同一天用同一头牛从两个牧场采集日粮样本。为了避免可能的牧场偏差,一周先从45-N牧场采集样本,一周后从0 - n牧场采集样本。Obioha等人(1970)报告了通过食道瘘采集的早晚饲料样本的氮水平有轻微但显著的差异。这种差异归因于饥饿导致的放牧选择性的差异。在我们的研究中,样本是在早上从牧场采集的,紧接着从第一个牧场采集样本。食管瘘的牛在邻近的O-N天然牧场上维持,不补充。在取样前一晚不给它们喂食,以便于收集饮食。每头牛收集的日粮样本在强制通风炉的浅平底锅中在50℃下干燥至恒定重量,然后在威利磨机中通过1毫米筛网研磨。化学分析包括凯氏定氮、体外可消化有机物(Tilley and Terry 1963, Moore and Mott 1974)、中性洗涤纤维(Van Soest and Wine 1967)、酸性洗涤纤维(ADF)和木质素(Van Soest and Wine 1968)。1978年和1979年,来自个体牛的饮食样本的处理方法与1977年相同。在每个样品上测量凯氏定氮,但在每个牧场的复合样品上进行其他化学分析。牧场管理杂志36(1),1963年1月。在夏季放牧季节,通过食道收集的草料样品的化学成分表明,从本地牧场接收45-N和O-N(1977年)。月IVDOM %牧场45-N设计CP NDF ADF D.M.木质素的% %的D.M. D.M. % % D.M.牧场的牧场放牧草场意味着45-N设计意味着45-N设计意味着45-N设计意味着45-N设计意味着可能“66.2 62.4 64.3 19.1 12.8 16.0 56.6 66.6 61.6 33.5 46.8 37.2 7.6 6.5 7.0 59.7 57.4 58.6 13.4 10.5 12.0 69.4 JO.6 6月7月70.0 39.1 44.3 41.7 6.4 8.2 7.3 56.6 53.6 55.1 12.7 8.8 10.8 10.6 JO.6 8月70.6 41.5 44.6 43.0 7.2 7.7 7.4 58.2 52.5 55.4 12.2 7.9 10.0 67.3 66.4 66.8 42.2 46.6 44.4 7.4 9.1 8.2 9月60.3 55.9 58.1 16.2 12.6 14.4 64.5 66I 65.3 39.1 46.8 43.0 6.8 9.6 8.2平均60.2 " 56.4 ' 58.3 14.7 " 10.5b 12.6 65.7 " 68磅66.9。1′44.6b 41.9′7.1′82 b 7.6′牧场同一品目不同上标牧场均值差异显著(K.05)。*每个值是20个样本的平均值(5只牛的4个日期),除了9月,只有3个采样日期,5月,45- n牧场有19个样本,0 - n牧场有8个样本。包含6月2日的抽样日期。每个抽样日期。数据按收集周期进行分析,采用单个饮食样本数据的算术平均值或复合样本的化学分析数据。在研究开始时便于补充的设备。1977年的研究20头体重约265公斤的一岁赫里福德阉牛被随机分配到45-N或O-N牧场。每个牧场的5头阉牛随机分配到以下处理:(1)不添加饲粮,或(2)9公斤干滚大麦(含2.7粗蛋白质和119克粗蛋白质)。在CDT前1000小时,每天早上将阉牛围在畜栏内,接受饲粮补充的阉牛单独饲喂。试验于6月1日开始,7月22日终止,原因是严重的干旱和随之而来的饲料缺乏,迫使阉牛离开牧场。所有的阉牛都连续使用50-50的盐和磷酸二钙混合物。分别对各牧场、各时期、各年份的增重情况进行分析,然后对2年ES数据和2年LS数据进行随机完整块分析。治疗差异由Duncan 's多重极差检验确定,在5%的概率水平上认为具有统计学显著性。
本文章由计算机程序翻译,如有差异,请以英文原文为准。
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