Spruce population under the canopy of overmature birch forests in the mixed forest zone of the european part of Russia

A. Deryugin, Yu.B. Glazunov, Yu.G. L’vov
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Abstract

The results of studies of the under-canopy population of spruce (Picea abies L.) in over-mature birch forests (Betuletum oxalidosum) of the Moscow region are presented. The purpose of the research was to study of the renewal process periodicity, age, vertical structure and condition of the spruce population. The work was carried out on a permanent trial plot in the period 2008–2018. The undulating feature of the spruce renewal process under the canopy of birch forest with a frequency of 30–40 years within the age of the birch trees under consideration has been established. It is determined that the basis of spruce forests formed under the canopy are trees with a renewal period of less than 30 years, which by the onset of the decay of birch forests form the first and second layers with an average age approaching the age of quantitative maturity. The age structure of the spruce population in birch forests aged 115 years is characterized by the dominance of trees of two age groups — 21–40 and 91–120 years, which account for 29 % and 37 % of the population, respectively. The emerging spruce forests belong to conditionally even-aged stands. Trees with a height of 2,0 m dominate (about 40 %) in the vertical structure of the population. The age dynamics of the stand is aimed at a slight increase in undergrowth trees and the first storey, and a decrease in the number of the second storey trees occurs. This is due to the emergence of new spruce specimens and the transition of part of the trees of the second storey to the first one. It was found that weakened trees predominate in the under-canopy population of spruce in overmature birch forests. Trees of the first layer are in the best condition (status category CS = 1). The state of undergrowth trees during the observation period did not change, remaining weakened (CS = 2,3–2,4). The set of trees of the second layer was characterized by positive dynamics — from the category of strongly weakened (CS = 2,7), due to the mortality, it moved to the category of weakened (CS = 2,0). Trees with a renewal period of up to 30 years are in the best condition. As a result of the succession process, after the decay of birch forests, low-density spruce forests with a productivity corresponding to the III site class will be formed.
俄罗斯欧洲部分混交林地带过熟桦树林冠下的云杉种群
本文介绍了莫斯科地区过成熟桦林(Betuletum oxalidosum)中云杉(Picea abies L.)林下种群的研究结果。本研究旨在研究云杉种群更新过程的周期性、年龄、垂直结构和状况。这项工作是在2008-2018年期间在一个永久性试验田进行的。建立了白桦林林冠下云杉更新过程的波动特征,其周期为30-40年。确定了林冠下形成的云杉林的基础是更新期小于30年的树木,而林冠下形成的第一层和第二层平均树龄接近于数量成熟期。115年龄桦林云杉种群的年龄结构表现为21 ~ 40年和91 ~ 120年两个年龄群的优势,分别占种群总数的29%和37%。新兴云杉林属于条件平均年龄林分。在种群的垂直结构中,高度为2000米的树木占主导地位(约占40%)。林分的年龄动态旨在下层树木和第一层树木的轻微增加,而第二层树木的数量减少。这是由于新的云杉标本的出现以及第二层部分树木向第一层的过渡。结果表明,过成熟桦林云杉冠下种群以弱乔木为主。第一层乔木处于最佳状态(状态类别CS = 1),观测期内林下乔木状态没有变化,保持较弱状态(CS = 2、3-2、4)。第二层树集的特征是正动态的——从强减弱的类别(CS = 2,7),由于死亡率,它移动到弱的类别(CS = 2,0)。更新期长达30年的树木处于最佳状态。在演替过程中,白桦林腐烂后,将形成生产力相当于III级立地级的低密度云杉林。
本文章由计算机程序翻译,如有差异,请以英文原文为准。
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