Tocharian B agent nouns in -ntsa and their origin

The agent noun suffix in -ntsa belongs to a complex of Tocharian B agent noun formations, similar in form, function, and inflection. Of these, two suffixes are widely believed to be related to -ntsa: the productive agent noun in -ñca and the lexicalised agent noun in -nta. The suffix -ntsa forms occupational titles to eleven verbs in Tocharian B and can be reconstructed for Proto-Tocharian through comparison with Tocharian A. In this paper, it is argued that the suffix originated in the feminine of the PIE active participle in *-nt. This is substantiated by the fact that several ntsa-nouns refer to primarily female professions, as well as the existence of the relic forms Bpreṃtsa ‘pregnant’ and Blāntsa ‘queen’. Furthermore, it is proposed that the masculine is reflected in the suffixes -ñca and -nta and that the disintegration of gendered inflection in the participle led to its development into several agent noun formations.