The late Pleistocene and Holocene history of Pinus koraiensis (Korean Pine) in the south of the Russian Far East based on palynological data

This paper reviews the distribution of Pinus koraiensis in the south of the Russian Far East in the past based on palynological and chronological data. The aim is to determine the chronological timing of the spatial migration of Pinus koraiensis and to correlate it with climate fluctuations explaining the changes of the geographical range of Pinus koraiensis in the south of the Russian Far East during the Marine Isotope Stages MIS 5 (127 000–71 000 BP), MIS 3 (57 000 BP – 28 000 cal BP), MIS 2 (28 000–11 700 cal BP), and MIS 1 (11 700 cal BP – to present). The consideration of the obtained pollen fossil data suggests that the range of Pinus koraiensis was in the south of the Russian Far East during MIS 5, as it was proved based on significant pollen grain percentage in the sediments of this age. Pinus koraiensis was widespread along with Pinus densiflora, Quercus, Ulmus, Juglans, Carpinus, and Phellodendron. During MIS 3, Pinus koraiensis was a component of birch-fir and birch-broadleaved forests. In MIS 2, Pinus koraiensis disappeared from the vegetation of this region. In the early Holocene, Pinus koraiensis expanded its range northward and again appeared in the vegetation of the south of the Russian Far East. Soon enough, it became one of the leading components in spruce-fir forests with Quercus and Ulmus. The increase in its proportion in the vegetation in the Holocene coincided mostly with the periods of warming climate from 11 700 to 11 500 cal BP, from 10 100 to 9300 cal BP, from 8800 to 5300 cal BP, from 4700 to 3500 cal BP, and from 2600 to 1800 cal BP. In the present case, the most diverse vegetation with Pinus koraiensis was typical of the Holocene optimum of the south of the Russian Far East from 8800 to 5300 cal BP.