Early-diverging Titanosauriform (Dinosauria, Sauropoda) Teeth from the Lower Cretaceous Yixian Formation of Southeastern Inner Mongolia, Northeast China

Three eusauropod teeth (SDUST-V1064, PMOL-AD00176, PMOL-ADt0005) are reported from the Lower Cretaceous Yixian Formation of Ningcheng, southeastern Inner Mongolia, China. Two of them (SDUST-V1064, PMOL-AD00176) are assigned to early-diverging titanosauriforms in having slightly mesiodistal expansion at the base of the tooth crown, a slenderness index value >2.0 and <4.0, and D-shaped cross section. Furthermore, SDUST-V1064 and PMOL-AD00176 are referred as an Euhelopus-like titanosauriform on the basis of having a sub-circular boss on the lingual surface and an asymmetrical crown-root margin which slants apically, respectively. CT scan data of SDUST-V1064 reveals new dental information of early-diverging titanosauriforms, for example, the enamel on the labial side thicker than that on the lingual side, an enamel/dentine ratio of 0.26 and a boss present on the lingual side of the dentine of the crown.