The Phylogenetic Relationships of Cretaceous Biting Midges, with a Key to All Known Genera (Diptera: Ceratopogonidae)

IF 1.1 3区 环境科学与生态学 Q3 BIODIVERSITY CONSERVATION
A. Borkent
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引用次数: 6

Abstract

ABSTRACT The phylogenetic positions of Cretaceous species of Ceratopogonidae previously placed in the genera Archiculicoides Szadziewski, Protoculicoides Boesel, and Atriculicoides Remm are reappraised in light of synapomorphies. Character states are discussed in detail, supported by new photographs of Protoculicoides depressus Boesel, the description of Protoculicoides revelatus, n. sp., from Burmese amber, and a compilation of previously published illustrations. The recent article by Szadziewski et al. (2016) proposing that Protoculicoides and Atriculicoides are congeneric is shown to be inaccurate. At least three separate lineages are represented by species in these two genera, requiring a new genus, Gerontodacus (type species, G. succineus (Szadziewski)), to include some of them. Archiculicoides, Protoculicoides, Gerontodacus, Adelohelea Borkent and Alautunmyia Borkent remain undetermined to subfamily. As a result of phylogenetic and other taxonomic considerations, the following are new combinations: Gerontodacus krzeminskii (Choufani, Azar, and Nel), Gerontodacus punctus (Borkent), Gerontodacus skalskii (Szadziewski and Arillo), Archiaustroconops andersoni (Szadziewski, Ross, and Giłka), Atriculicoides ciliatus (Borkent), Atriculicoides hispanicus (Szadziewski and Arillo), Atriculicoides sanjusti (Szadziewski and Arillo) and Adelohelea burmitica (Szadziewski and Poinar). The following species are returned to the genera they were assigned to before Szadziewski et al. (2016): Atriculicoides cenomanensis Szadziewski and Schlüter, Atriculicoides dasyheleis Szadziewski, Atriculicoides globosus (Boesel), Atriculicoides incompletus Szadziewski and Schlüter, Atriculicoides macrophthalmus Remm, Atriculicoides sibiricus Szadziewski, Atriculicoides swinhoei (Cockerell), Atriculicoides szadziewskii Pérez-de la Fuente, Delclòs, Peñalver, and Arillo and Atriculicoides taimyricus Szadziewski. A key is provided to all Cretaceous Ceratopogonidae genera.
白垩纪蠓类的系统发育关系(双翅目:蠓科)
摘要对白垩纪蠓科原属Archiculicoides Szadziewski属、Protoculicoides Boesel属和atricicoides Remm属的蠓科物种的系统发育地位进行了重新评价。本文通过对Protoculicoides depressus Boesel的新照片、对缅甸琥珀中Protoculicoides revelatus, n. sp的描述以及先前发表的插图汇编,详细讨论了其特征状态。Szadziewski等人(2016)最近的一篇文章提出Protoculicoides和atricicoides是同源的,这被证明是不准确的。这两个属中的种至少代表了三个独立的谱系,需要一个新的属,Gerontodacus(模式种,G. succineus (Szadziewski)),以包括其中的一些。Archiculicoides, Protoculicoides, Gerontodacus, Adelohelea Borkent和Alautunmyia Borkent的亚科尚未确定。以下物种回归到Szadziewski et al.(2016)之前归属的属:萨曼蠓和schl、达西蠓、球形蠓、不完全蠓和schl、大眼蠓、西伯利亚蠓、斯文蠓、萨德兹耶夫斯基蠓、萨德兹耶夫斯基蠓、萨德兹耶夫斯基蠓、萨德兹耶夫斯基蠓、Delclòs、Peñalver以及萨德兹耶夫斯基蠓和泰米斯基蠓。提供了白垩纪所有蠓科属的钥匙。
本文章由计算机程序翻译,如有差异,请以英文原文为准。
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来源期刊
American Museum Novitates
American Museum Novitates 环境科学-动物学
CiteScore
3.00
自引率
6.70%
发文量
8
审稿时长
>36 weeks
期刊介绍: The Novitates (Latin for "new acquaintances"), published continuously and numbered consecutively since 1921, are short papers that contain descriptions of new forms and reports in zoology, paleontology, and geology.
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