The Terminal Assemblage of Permian Brachiopods in the Verkhoyanie, Northeast Russia

Pub Date : 2024-04-06 DOI:10.1134/s1819714024010068
R. V. Kutygin, V. I. Makoshin, A. S. Biakov
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Abstract

Brachiopods from the upper part of the Khalpirkian Regional Stage of the Tiryakh–Kobyume section of the Kobyume structural–facies zone in Southern Verkhoyanie have been studied. The collection is dominated by Strophalosia? vollossovitschi (Fredericks) and Cancrinella? sp. Single spiriferids are assigned to Crassispirifer monumentalis Abramov et Grigorjeva and Cr. aff. monumentalis Abramov et Grigorjeva, and, thus, host deposits are referred to the Crassispirifer monumentalis Beds. We found these beds earlier only in the lower part of the Khalpirkian Regional Stage. Reliable findings of the terminal brachiopod assemblage confined to the Khalpirkian of the Upper Permian have been reviewed. The assemblage is dominated by strophalosiids represented by Wyndhamia gijigensis (Zavodowsky), Marginalosia? magna Abramov et Grigorjeva, Subtaeniothaerus lungersgauzeni Solomina, and Strophalosia? aff. vollossovitschi (Fredericks). “Magadania” sp. and Cancrinella? sp. have been identified from linoproductids. Crassispirifer monumentalis spiriferids are an important element of the terminal assemblage. Athyridids of the Khalpirkian Regional Stage, known only in its lower part, are represented by Cleiothyridina nikolaevi Grunt, Cleiothyridina sp., and Bajtugania sp. The studied regional stage also contains findings of terebratulids attributed to Marinurnula? aff. mantuanensis (Campbell) and M.? aff. chivatschense (Zavodowsky). No brachiopods are known at the top of the Khalpirkian Regional Stage, while rhynchonellids Piarorhynchia sp. appear at the base of the Nekuchan Regional Stage. At the beginning of the Khalpirkian, the brachiopod community had an extended natural habitat in the shelf zone of the Verkhoyan Sea along the Angaraland coastline. Subsequently, this area was reduced and initially divided into two independent subareas confined to the East Kharaulakh and South Verkhoyan sectors. In the Late Khalpirkian, the brachiopod community was concentrated only in the Kobyume zone, where it ended up under the end-Permian mass extinction. Some taxa of the identified brachiopod assemblage have been traced in the Okhotsk region, the Ayan—Yuryakh anticlinorium, and the Omolon massif.

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俄罗斯东北部 Verkhoyanie 地区二叠纪腕足动物的末期组合
摘要 研究了南上甘岭科比乌梅构造成因区 Tiryakh-Kobyume 段 Khalpirkian 地区阶段上部的腕足类动物。采集到的主要是Strophalosia? vollossovitschi(弗雷德里克斯)和Cancrinella? sp.单螺旋体被归入Crassispirifer monumentalis Abramov et Grigorjeva和Cr.我们早先只在 Khalpirkian 地区层的下部发现了这些岩床。对仅限于上二叠统 Khalpirkian 的末端腕足动物群的可靠发现进行了回顾。腕足动物群主要由石龙子类代表,包括 Wyndhamia gijigensis (Zavodowsky), Marginalosia? magna Abramov et Grigorjeva, Subtaeniothaerus lungersgauzeni Solomina, 以及 Strophalosia? aff."Magadania" sp.和 Cancrinella?Crassispirifer monumentalis螺旋体是末端集合体的重要组成部分。Khalpirkian 地区阶段的甲虫类仅在其下部有所了解,代表有 Cleiothyridina nikolaevi Grunt、Cleiothyridina sp.和 Bajtugania sp.。 在所研究的地区阶段还发现了蝶形虫,分别属于 Marinurnula? aff. mantuanensis (Campbell) 和 M.? aff. chivatschense (Zavodowsky)。在 Khalpirkian 地区阶段的顶部没有发现腕足动物,而在 Nekuchan 地区阶段的底部则出现了菱形纲的 Piarorhynchia sp.。在 Khalpirkian 早期,腕足动物群落在安加拉兰海岸线沿线的上霍延海陆架区拥有广阔的自然栖息地。随后,这一区域逐渐缩小,最初被划分为两个独立的分区,分别局限于东哈拉乌拉克区和南维尔霍扬区。在晚 Khalpirkian 期,腕足动物群落仅集中在 Kobyume 区,并在那里经历了二叠纪末的大灭绝。在鄂霍次克地区、阿扬-尤里亚克反褶带和奥莫隆地块,已经发现了腕足动物群的一些分类群。
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