Nepenthes (Nepenthaceae) of Halmahera, Indonesia

Two new paniculate species of Nepenthes, N. halmahera and N. weda, both allied to N. danseri Jebb & Cheek, are described respectively from lowland and lower montane forest on ultramafic substrate. Nepenthes weda appears to be unique in the genus due to the adaxial tepal surfaces which in the distal portion are hairy and lack nectar glands. This species is also unique among paniculate members of the genus in its forward-facing, concave subapical lid appendage. Both species are currently only known from the Weda Bay Nickel Project concession area in Halmahera, Indonesia, and are assessed as Critically Endangered using the 2012 IUCN standard. Two character states are formally described and named for the first time in Nepenthes: nanophyll rosettes and multiseriate fringed pitcher wings. Stage-related heteromorphy in lid appendages is documented for the first time in the genus. Keys to the species of Nepenthes of Halmahera, and to the paniculate species of SE Asia are presented.