Furongian (upper Cambrian) trilobites and agnostoids from the Alum Shale Formation of Bornholm, Denmark: revised taxonomy and biostratigraphy

The Furongian (upper Cambrian) trilobite-agnostoid fauna from the Alum Shale Formation of Bornholm, Denmark, is reviewed and revised. The study is based on the museum material stored at the Natural History Museum of Denmark, including the material originally monographed by C. Poulsen (1923) [Bornholms Olenuslag og deres fauna. Danmarks Geologiske Undersøgelse II. Række, Vol. 40, 83 pp]. A total of 8502 specimens, mostly disarticulated sclerites, have been registered. The taxonomy of all species is updated and the best preserved specimens are illustrated. A total of 39 olenid and 5 agnostoid taxa (incl. the Miaolingian Agnostus pisiformis) are recorded including one new species, Ctenopyge magna n. sp. Two specimens of Ctenopyge, treated in open nomenclature as Ctenopyge sp. 1 and sp. 2, may also rep-resent new species. 14 taxa have not been previously reported from Bornholm, viz. Ctenopyge ahlbergi, Ctenopyge tumidoides, Eurycare brevicauda, Leptoplastus abnormis, Leptoplastus crassicornis, Olenus transversus, Parabolina lobata praecurrens, Peltura acu-tidens, Peltura minor, Peltura westergaardi, Protopeltura planicauda, Protopeltura praecursor, Pseudagnostus leptoplastorum? and Sphaerophthalmus drytonensis. Ctenopyge pecten and Ctenopyge affinis are also new to Bornholm as the material formerly described under these names represent Ctenopyge tenuis and C. magna n. sp., respectively. Lotagnostus americanus, Ctenopyge fletcheri, Sphaerophthalmus alatus and Triangulopyge humilis were described under different names by C. Poulsen (1923). Peltura westergaardi and Ctenopyge tenuis are elevated from subspecies to species rank. A redescription of Leptoplastus bornholmensis is presented; the species is transferred to Eurycare. The identification of isolated skeletal parts of L. abnormis and Leptoplastus ovatus and Sphaerophthalmus flagellifer and S. drytonensis are remarked on. The presence of three agnostoid and 14 trilobite zones is confirmed by fossils and all six Furongian superzones are developed on Bornholm. At least the Leptoplastus paucisegmentatus and Leptoplastus raphidophorus zones seem to be absent. Other undocumented zones may be unfossiliferous, not exposed or truly absent. Three different trilobite assemblages (potential subzones) are discerned in the Peltura acu-tidens–Ctenopyge tumida Zone; Ctenopyge tumidoides and Sphaerophthalmus angustus range into the basal part of this zone. All exposures of the Furongian Alum Shale Formation along the Læså and Øleå streams on southern Bornholm are briefly described including GPS coordinates.