A new Albian genus and species and two other new species of Umenocoleidae (Dictyoptera) from South Korea

Since the first description from China, the Mesozoic dictyopteran family Umenocoleidae was recorded from the Eurasian and South American continents with 18 genera and 31 species. Here, we describe one new genus and three species from the Jinju Formation, South Korea: Umenocoleus minimus sp. nov., Pseudoblattapterix weoni gen. et sp. nov., and Petropterix koreaensis sp. nov. We also discuss three new apomorphies of the Umenocoleidae and Alienopteridae, which are 1) M, Cu and PCu1 veins form an common vein near wing base, 2) M, Cu and PCu1 veins appress to R vein and 3) CuP + PCu1 vein are not as concave as in the other Holopandictyoptera (total group of extant Blattodea and Mantodea). On the basis of these new apomorphies, we support the inclusion of the family Alienopteridae into the superfamily Umenocoleoidea. We also support the exclusions of the genera Vitisma and Permoponopterix from the Umenocoleidae family and propose to transfer the genus Permoponopterix into the Protelytroptera.